i9o8] EDGERTON—ANTHRACNOSES 385 



ment, and these have been described during the past few years for a 

 number of forms^ by a number of investigators. Only a few things 

 which do not seem to have been made clear need to be mentioned. 

 The conidia germinate very readily in water or in nutrient solu- 

 tion, generally w^ithin three or four hours. The germ tube may be 

 sent out from any place on the spore, but in most cases the first tube 

 originates near one end. It rarely comes exactly from the end, but 

 very close to it, so that it appears to extend at an angle to the main 

 axis of the spore {iigs. 30 ^ j6y j/). If it came exactly from the end, 



I 



the germ tube and the spore would lie in a straight line. In some cases 

 the tube does appear to come from the end, but this seems to be un- 

 common as compared with the other method. After the first germ 



may develop apparently from any place on the 



spore. 



Germination differs somewhat in different media. Rarely in 

 nutrient solutions does the spore become septate in germination. In 



more 



much 



the 



spore becomes septate. . This variation in germination in different 



seems 



to be characteristic of all members of this type. It is 



+ 



interesting at this point to note a recent paper by Delacroix (12) on 

 Gloeospormm nmsarum. In studying the germination of its spores, 

 he found that* they nearly always became septate. Miss Stoneman 

 (43} some years previously, in w^orking on the same form, had figured 

 the spores as continuous. Delacroix was undecided whether to 

 call the form he was w^orking with a new species or not, as this was 

 the only point of difference. If he had germinated his spores on 

 different media, he imdoubtedly would have found both types of 

 germination. 



If germination takes place in a medium lacking nutrient material, 

 as water, the germ tubes grow only a short distance and then form 

 at their tips rather large brown cells more or less variable in shape 

 ^S^- 3^, 33)- These have been spoken of variously by different 

 investigators as secondary spores, appressoria (Hasselbring 24), 

 etc. They germinate readily under the right conditions, although 

 some seem to have had difl&culty in obtaining germination. If nutri- 



o 



mi 



may be formed at the end of this germ tube (fig 



