1908] EDGERTON—ANTHRACNOSES 403 



this time it has gone without question that these forms are all the 

 same. Spaulding and von Schrenk (33) state that they have seen 

 specimens of the bitter rot op apples in Vermont as late as October. 

 They might have seen apples affected with a Gloeosporlum^ but it 

 is not certain that the apples were affected with the same form that 

 grows in the south- As was stated above, the two form? on culture 

 media are very distinct, and on the apple itself there is a tendency to 

 divergence. On the apple from the south the conidial pustule as a 

 rule is large^ containing much pseudo-parenchymatous tissue which 

 protrudes from the surface {jig. 12) ; while the northern form produces 

 a pustule with little pseudo-parenchymatous tissue, similar to that 

 illustrated from the tomato {fig. ii). This distinction does not 

 always separate the two forms, as they often intergrade, but the 

 characters are noticeable in studving the two. Furthermore, the 



form 



while the northern 



form, so far as is known, never produces them. If we call these two 



mi 



sporium into one species and say that Glomerella is monotypic, grow- 

 ing on nearly everything. But if we call them distinct, what shall 

 we name the forms ? The names rujomaculans and Jruciigemim were 

 given by Berkeley (6, 7) to forms in England on grape and apple. 

 These were found in a latitude comparable to our northern states, 

 and consequently we should think that Berkeley had a form com- 

 parable to our northern one. If this is true, the perfect stage of the 



Gloeosporitim ruf> 



This north- 



ern form is known to occur on several other hosts, not entirely fruits, 

 besides the apple. The anthracnosc of tomatoes has generally been 

 called Gioeosporiiim phomoides, and under this name it has been 

 discussed by several authors. But the culture obtained by me was 

 absolutely identical with the northern form on apple; although the 



Mi 



culture from apple. 



ph 



moides takes exception to Miss Stoneman's conception of the species. 

 Tt seems that G. phomoides is an entirely different thing, and does not 

 belong to the Glomerella tvDe of anthracnoses; hence the name can 

 no longer be used for the anthracnose commonly seen on tomatoes. 

 The forms from grape stems, raspberry stems, and Podophyllum 



