348 BOTANICAL GAZETTE [may 



lower series occurs, as described above. All of the upper series of 

 bundles pass into the base of the ovule, the only xylem element^ 

 found beyond the ovule in this region being a new development asso- 

 ciated with the main bundle. A single resin duct accompanies the 

 latter throughout its course. This does not branch and is the only 

 representative of the tissue in the scale, no canals being formed in 

 connection with the upper series of bundles. 4 



The vascular system of the axis of the megasporangiate cone con- 

 sists of a ring of collateral bundles very similar in general to that of 

 the microsporangiate cone {fig. 8) . The wood of these is usually of 

 the ordinary endarch type, but near the base of the cone there are 

 often a few centripetal elements associated with the bundles at the 

 sides of the gap left by the exit of the megasporophyll trace. In one 

 instance these were almost in continuity with the wood elements of the 

 ovular supply in the sporophyll. The latter, especially in its young 

 Condition, appears very much of the nature of centripetal xylem 

 (fig. 12),* and the occurrence of the isolated elements in the axis 

 affords confirmation of this idea. They are found also in the upper 

 part of the pedicel, but in my material they are rare here. 



The scales at the base of the micro- and megasporangiate cones and 

 the foliage leaves both receive a single vascular bundle and a sing e 

 resin duct from the axis, the supply coming off in a similar way to t at 

 of the sporophylls. There is always a gap in the cylindrical ste e 

 opposite the "trace" to each of these members {fig. 8, for microsporo- 

 phylls, lower right hand side, etc.) . . 



In several of the megasporangiate cones of Saxegothaea a ev\ o 

 the lower sporophylls bear ovules on their under surface (Jig- >' 

 These have an integument, but not the epimatium or rudimen ar> 

 second integument of the normal ovule. They stand out too ro^ 

 the sporophyll more freely, and are further developed than the upp^ 

 ones in the same cone. A slight vascular supply goes off to same 

 these, the bast and wood showing a tendency to orient itself b _. 

 to that of the main bundle of the scale. This supply passes aro 

 the resin duct. 



*Cf. Stiles, Lc. 216. mature 



s Miss Stiles has come to a similar conclusion from studying the mo 

 condition. 



