1918] Yocum: The Neuromotor Apparatus of Euplotes Patella 345 



from the posterior ends of five longitudinal grooves over the posterior 

 margin of the body. Along the posterior margin of the body are four 

 small marginal cirri. The marginal cirrus to the right is always 

 fimbricated, as is usually the one next to it. 



The five furrows in which the anal cirri lie are formed by slight 

 indentations of the pellicle along the line of the furrow and also a 

 slight uprising of the pellicle between the furrows. These ridges are 

 just far enough apart to permit of the insertion of the bases of the 

 cirri. Such a close arrangement permits the anal cirri to move in only 

 one plane, that parallel to the median plane of the body. Of the six 

 ridges bounding the five furrows, if we number from right to left, the 

 first, fourth, and sixth are almost twice as long as the other three, and 

 extend from the cirri toward the anterior right corner of the cytostome 

 for fifty to sixty microns. All stained specimens show a distinct granu- 

 lation of the posterior ten to fifteen microns of the ectoplasm of the 

 ridges. The granules of these areas are much smaller than those in 

 the other parts of the ectoplasm and when stained give the posterior 

 parts of the ridges a clouded appearance. Such a granulation near the 

 bases of the cirri indicates that a possible physiological relationship 

 exists betwen the cirri and adjacent ectoplasm. 



Swezy (1916) pointed out that the parabasal body of certain simple 

 flagellates, such as Prowazekia lacertae, begins as a finely granular 

 chromidial body lying near the basal granule or blepharoplast of the 

 flagella. The function of the parabasal body is interpreted as a kinetic 

 reservoir or a body of reserve kinetic material. Usually this para- 

 basal body is connected to the blepharoplast by a rhizopla.st. "While 

 in Euplotes patella no definite threadlike connection has been found to 

 exist between the granular portions of the ridges and the cirri between 

 them, it seems probable that a relationship exists between them com- 

 parable to that between the parabasal body and the blepharoplast of 

 flagellates. If this be true the granules are to be considered as indi- 

 cative of reserve material which by oxidation liberates energy neces- 

 sary for the vigorous movements of the cirri. While not so prominent, 

 it is noticeable that around the bases of all of the cirri on the animal 

 there is an area about one micron wide which retains the haematoxylin 

 dyes to a greater degree than the main body of the cytoplasm. 



At the anterior end of the animal is a crescent-shaped lip about 

 four microns wide at its middle point. In his discussion of Euplotes 

 patella, Stein (1859) describes the blunt anterior end of the animal as 

 terminating in a three-cornered lip. In no case have I seen this lip 



