Species Diversity 



Mean annual species diversity (H') ranged from 2.6 to 4.6 (Table 7). Low diversity indices (e.g., 

 1980 at F,F and 1984 at JC) were seen when short-term pulses in species abundance caused the evenness 

 component of diversity to decline. High H' values reflected changes in the numbers of individuals or 

 species, but not in the equality of the distribution of individuals among species. 



0.1 3.8+0.1 3.9+0.1 3.4+0.3 3.4+0.2 3.8+0.3 

 0.1 0.7 + 0.1 0.7 + 0.1 0.7 + 0.1 0.7 + 0.1 0.7 + 0.1 



445 + 335 907 + 293 



39 



