202 BOTANICAL GAZETTE [SEPTEMBER 
The antheridium rudiment is elongated, and it divides transversely 
into six or seven primary cells. The wall cells and spermatogenous 
cells are separated from each other in the body of the antheridium 
after the formation of quadrant and octant walls. The mature 
antheridium is elongated and pointed and is sunken in the receptacle. 
In the nucleus of the spermatozoid the individual chromosomes 
are recognizable as distinct twisted fibers. 
The archegonium is very long-necked, has six rows of neck cells 
and twelve or more neck canal cells. It is probably fertilized before 
the hood-like involucre has grown far beyond its tip. 
The capsular portion of the sporogonium divides to quadrants 
and octants before sporogenous cells and wall cells are separated. 
The foot is small, the seta stretches to 30 or 40™™ in length, and the 
extended capsule is erect, elongated, cylindrical, and its wall is a 
single layer of cells. 
Monoclea possesses two kinds of rhizoids, corresponding to those of 
Marchantia in size, direction of growth, and in the presence of 
tubercles in those of one type. 
The absence of air chambers and ventral scales is probably due 
to the nearly aquatic habit of the plant. 
The evidence gained from the study of the origin and structure of 
the male receptacle, and of the antheridium and archegonium, and 
from the structure of the wall of the capsule, and the presen and 
direction of growth of the two types of rhizoids, favors the view that 
Monoclea is most closely related to the lower Marchantiaceae. 
Jouns Hopkins UNIVERSITY, 
Baltimore, Maryland. 
LITERATURE CITED. ; 
Biscuorr, G. W., Bemerkungen ueber die Lebermoose, vorziiglich aus . 
Gruppen Marchantieen und Riccieen, etc. Nov. Act. Acad. Caes. Leop. 
Car. 177: gog-1088. pls. 67-71. 1835. 
CAMPBELL, D. H., Mosses and Ferns. 1895. ‘7 
, The systematic position of the genus Monoclea. Bor. GAZETTE sap 
274. 1898. 
Cavers, F -, On the structure and biology of Fegatella conica. Annals of itd 
18:87-120. pls. 6-7. 1904. «298. gs 
Coxer, W. C., Selected notes, II, Liverworts. Bor. GAZETIE i 
I-5. 1903. 
