oa: BRIEFER ARTICLES 301 
With the inception of zonation comes the advance from the spirem to 
the later stages of mitotic division in all the nuclei of the oogonium. Many 
of the nuclei in process of division are floated to the periplasm, others com- 
plete the division within the ooplasm and there leave both daughter nuclei 
within the egg. The completion of this mitosis results in conditions very 
like those in A. Tragopogonis.s Many of the nuclei are in the periplasm, 
some are in the ooplasm. The chief difference lies in the coenocentrum, 
to be discussed later. 
The daughter nuclei of the first division, which remain within the 
ooplasm, proceed to a second mitosis as in A. Tragopogonis and A. candida, 
though the number of oospheric nuclei is constantly diminishing, owing 
lo their outward movement. All of them, except one or two contiguous 
to the coenocentrum, reach the periplasm before or immediately after 
mitosis is completed. One or two nuclei in mitosis are seen attached to 
the coenocentrum during the second division. The completion of the 
mitosis, however, finds only one daughter nucleus remaining thus to func- 
ton as the female pronucleus. Conclusive evidence was not obtained, 
but itis probable that some nuclei suffer degeneration within the ooplasm, 
asis the rule in A, Tragopogonis. 
; The coenocentrum is first to be seen as a homogeneous globule, much 
like the central globule of A. Bliti, though considerably larger, being nearly 
as large as the nuclei at the completion of the first mitosis. There is no 
striking differentiation of the protoplasm surrounding the globule at this 
ee The globule, which does not change materially in size in later 
cae . me than that of A. Bliti and much smaller than that of A. 
na ms : shows much less structural differentiation than is exhibited 
a ule in the latter species. When the second mitosis approaches 
phase the globule is surrounded by zones of protoplasm of varying 
ne . i nee resemble the structure shown in Ags. 69 and 
7 eg - Bliti,® though the development attains a higher degree 
~panduranae. 
The attractive 
attachment of 
(igs. x » 2), 
power of the coenocentrum for nuclei is evidenced by the 
, Shorter in this than in most species of Albugo, 
of the completion of the second division, emptying 
"ENS, F. 1... Joc. c4., pl. 3, fig. Bt. 
® Steven 
pie The compound oosphere of Albugo Bliti. loc. cit. pl. 13. 
