1905] COULTER & LAND~—TORREYA TAXIFOLIA 175 
showed that precisely the same explanation applies to them that we 
have given in the case of Torreya. 
SUMMARY. 
The staminate strobilus consists of a series of closely overlapping 
sterile bracts, in four vertical rows, completely enveloping the tip of 
the axis bearing numerous stamens. The large adaxial resin cavity 
that occurs in the stamen occupies the site of three abortive sporangia. 
The male gametophyte has no prothallial cell, and the male cells 
are very unequal, resembling those of Taxus. The pollen tube is 
exceedingly variable in the rate and direction of its advance through 
the nucellar cap, sometimes pushing in the embryo sac while it is 
in an early free-nucleate stage. 
The ovulate strobilus consists of four enveloping bracts and a 
single terminal ovule with two integuments. Extensive intercalary 
growth below the mother-cell forms the bulk of the mature ovule 
and seed. There is no organization of a special digestive layer 
around the mother-cell. 
The solitary archegonium initial appears as soon as walls are 
formed, is always at one side of the central axis of the gametophyte, 
and forms a two-celled neck. The nucleus of the central cell was 
not observed to divide, nor could any trace of a ventral nucleus be 
found. 
In fertilization the male cytoplasm invests the fusion nucleus, 
and seems to remain distinct until wall-formation at the four- 
nucleate stage of the proembryo. 
In the development of the proembryo, four free nuclei appear 
before wall-formation, and the proembryo completely fills the egg, 
having no “open cells.” A proembryo of twelve to eighteen cells 
is the winter stage. In the spring the suspensor is formed by what 
may be called a wave of elongation, beginning with the uppermost 
tier of the proembryo, and extending gradually downward, tier after 
tier, until it includes the upper region of the meristematic cylinder 
formed by the terminal cell. 
Small embryos are formed during the second season in the sus- 
pensor region of the normal embryo; but whether they arise from 
Prothallial or suspensor cells was not determined. 
