ON EXPERIMENTAL STUDIES IN THE PHYSIOLOGY OF HEREDITY. 247 
be inherited through the egg-cell only, the male gamete playing no part 
in determining the nature of the offspring in respect of these characters. 
The experiments of Mr. R. P. Gregory and Miss Gairdner on the 
inheritance of variegation and other characters in Trope@olum have been 
continued. It is hoped that sufficient data will have been gained by 
the end of the present season to permit of the publication of an account 
of this work. Present results indicate that in Trop@olum variegation 
is inherited in the usual way from both the father and the mother, and 
is a Mendelian recessive character. Other characters in Tropeolum 
which are being studied are those of colour and habit (dwarf or trailing). 
The experiments on the gynandrous variety of the Wallflower and 
its relation to the normal type are nearing completion, and it is hoped 
that an account of them will be published next year. 
The Committee ask for reappointment with a grant of 45]. The 
expenses of these experiments involve an annual outlay of about 1101. 
to 1201. By far the largest item in this expenditure is the cost of 
labour, which has increased during the last few years with the general 
rise in wages which has taken place. Other important items are those 
of the rent of the garden and the cost of heating the Primula house. 
During the present year Miss Saunders and Mr. Gregory jointly receive 
a grant from the Royal Society of 601. in aid.of the cost of this work. 
Breeding Experiments with Ginotheras.—Report of the Com- 
mittee, consisting of Professor W. BATEson (Chairman), Pro- 
fessor F. KEEBLE (Secretary), and Mr. R. P. GReEcory, 
appointed to carry out the Experiments. 
Tur Committee have received the following Report from Dr. R. R. 
Gates on the experiments which he has made :— 
‘The grant of 201. made by the British Association for Gnothera- 
breeding has been applied to the expenses of these experiments during 
the last year. In the season of 1913 about 10,000 plants were grown, 
representing a great many races and hybrids of Ginothera. The plants 
were grown at Rothamsted on a two-acre plot set apart for the purpose. 
They developed very successfully, nearly every individual reaching 
maturity. The largest series of hybrids were the F, from @. grandi- 
flora, GH. rubricalyx and its reciprocal, and the F, of crosses between 
CH. grandiflora and Gi. Lamarckiana. The F, generation of the former 
cross confirms and extends the results of the F, and F, generations 
already published in ‘ Zeitschr. f. Abst. u. Vererb.,’ vol. xi. They 
show in particular that both blending and alternative inheritance of 
characters occur. Some of the plants, which have been examined 
eytologically in conjunction with Miss Nesta Thomas, further emphasise 
the fact that mutation and hybridisation in Cnothera are separate 
processes, both of which may go on together. Some of these results 
will be incorporated in a book now in preparation.’ 
