Bourne—On the Post-Embryonic Development of Fungia. 246 
The Caryophyllia form of the Trophozooid or Anthoblast is 
not long maintained, but the margin ofthe calyx grows outwards 
to form a disc-shaped Anthocyathus. This is eventually set free, 
and new Anthocyathi, to the number of three or four, are succes- 
sively formed and detached from the Anthocaulus. In a young 
Trophozooid or Anthoblast there are twelve primary septa; in 
succeeding stages—twelve secondaries, twenty-four tertiaries, and 
forty-eight quaternaries are developed. The relations of the ter- 
tiaries to the secondaries and the quaternaries to the tertiaries are 
the same as in Stephanophyllia. Fungia cannot be placed in any 
of the groups Huthecalia, Pseudothecalia, or Athecalia, established 
by recent authors. A compact theca is present, formed partly by 
the union of the peripheral ends of the septa, partly by interstitial 
pieces uniting the peripheral ends of the septa. ‘True epitheca is 
present in the Anthoblast. 
The anatomy of the polype resembles that of Actinia. Typi- 
cally each septum is enclosed by a pair of mesenteries; but, as in 
course of growth a new cycle of septa makes its appearance before 
the corresponding mesenteries, there are stages in which the septa 
are—(qa) all entoccelic, (0) alternately exocolic and entoccelic. 
The structures of the Anthocyathus are formed as direct out- 
growths ofthe corresponding. structures in the Anthocaulus; but 
the mesenteries are scarcely represented in the Anthocaulus, and, 
after detachment of an Anthocyathus, are formed anew in the 
distal end of the Anthocaulus before the outgrowth of a new 
Anthocyathus is externally visible. 
The absorption of the corallum at the point where the Antho- 
caulus passes into the Anthocyathus, causing the detachment of 
the latter, is not effected by means of phagocytes, nor through the 
agency of a special tract of tissue cells. Numerous dark-staining 
bodies of peculiar character have been observed, but they occur in 
all parts of the Anthocyathus, and in many parts of the Antho- 
caulus, and, probably, are nutrient cells, and are notin any way 
connected with the absorption of the corallum. In the region 
where absorption of the corallum, and consequently the detach- 
ment of the Anthocyathus, takes place, the tissues undergo dege- 
neration, and the corallum contiguous to the degenerate tissue 
becomes white, opaque, and brittle. The parts of the corallum 
which are thus altered are especially liable to the attacks of the 
