Lkonard — Genus Taenitis, "with Notes on remaining Taenitidinae. 263 



The venation of Eschatoi/ramme is similar to that in Taenitis, and unlike 

 the remaining form in the absence of intra-areolar vein-endings. The reticu- 

 lation is rather meagre, consisting chiefly of a series of large areolae on either 

 side of the midrib, while free vein-endings are directed outwards towards the 

 margin. It might readily be. held as reduced from the condition seen in 

 Taenitis (Fig. 3, a and b). 



Diels ((5), p. 303) says that the veins of the sterile leaves of Dicrano- 

 glossum (= Eschatogramme) are mostly free and seldom anastomose. It will 

 be seen from the drawings of the sterile leaves (-Fig. 6, a and b) that, on the 

 other hand, the veins are mostly anastomosed, and seldom free. In the fertile 

 leaves, there is a very definite commissure beneath the sorus (Fig. 6 c, and (5), 

 p. 301). This is another point of resemblance to Taenitis and Blcchnum. 



The reticulation of the veins, and the presence of a soral commissure, are 

 close links to Taenitis, and it seems probable therefore that there is aphyletic 

 sequence from Tucnitis to Eschatogramme, and on to forms in which the 

 pinnation of the leaf has completely disappeared, the result being the simple 

 leaf. 



The sporangia are very similar to those of Taenitis. There is an inter- 

 rupted annulus with a variable number of indurated cells — 12 to 18 The 

 stalk is a three-celled one, and the spores are thin-walled and two-sided. 



Drymoglossum. 



Drymoglossum was first described by Presl in 1836. It has gone under 

 various synonyms, amongst them being Taenitis, in which genus it was placed 

 by Mettenius in 1856. It habit is seen in (5), fig. 160. It has a slender 

 elongated rhizome which is covered with scales. The leaves which are 

 dimorphic arise some distance apart from each other. The fertile leaves are 

 long and linear, while the sterile are small and ovate. The sorus is con- 

 tinuous and without a covering. There are nine species, of which D. camosum 

 and D. hetcrophyllum' were available for examination. 



Dermal Appendages. 



The dermal appendages of Drymoglossum are scales only (Fig. 7, a). They 

 are broad and well-developed, and are attached to the epidermis at approxi- 

 mately their centre (x). They form a striking contrast to the small hair-like 

 scales of Taenitis, and this is probably in relation to its epiphytic habit. 



Anatomy of Stem and Leaf. . 

 The stelar anatomy of Drymoglossum shows an advanced dictyostelic con- 

 dition. The number of meristeles present vary from four to seven, and there 



SCIENT. PKOC. R.D.S.,VOL. XV., NO. XXIV. 2 U 



