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ON OENOTHERA, ETC. 421 
of Anthropology, but the Munro Lecturer gives ten lectures each year on 
Anthropology or Prehistoric Archeology. 
The Royal Scottish Museum and the Scottish National Museum of Antiquities 
have important collections of Scottish and other antiquities, and the Museum 
of the Royal College of Surgeons a large collection of anatomical material. 
Glasgow.—The University has a fair teaching collection for Physical Anthro- 
pology with about 300 skulls, representing most of the racial groups. The 
Hunterian Museum contains the considerable Turner collection of ethno- 
graphical material from the South Seas; a representative collection of Scottish 
antiquities, prehistoric and Roman; an important collection of ancient coins, 
and other classical and Egyptian material. The Library is open to matriculated 
students, and research students also may be admitted. Laboratory accommoda- 
tion for research students is provided as required. 
The Professor of Anatomy, Dr. T. H. Bryce, F.R.S., gives instruction in 
Physical Anthropology, and the Hon. Curator of the archeological and ethno- 
graphical collections of the Hunterian Museum. There is an independent 
Department of Geography, under a Lecturer; and instruction in Classical 
Archeology is given in the Departments of Greek and Latin. 
Except in Physical Anthropology, no ordinary degree or diploma of the 
University includes anthropological studies; but a research student may 
present a thesis in any branch of Anthropology for the degree of Ph.D. after 
three years’ research in the University or in a College affiliated thereto. 
Oenothera, &c.— Final Report of Committee (Dr. A. B. Renpxe, 
Chairman; Prof. R. R. Gates, Secretary; Prof. W. Bateson, 
Dr. W. Brieruey, Prof. O. V. DarpisHire, Dr. M. C. Rayner) 
appointed to continue Breeding Experiments on Oenothera and 
other Genera. 
Correlated cytological and genetical experiments with a number of genera 
have been continued at the Royal Botanic Gardens, Regent’s Park, during the 
past year. These genera include Oenothera, Celosia, Lathyrus, Brassica, Tiarella, 
and Lactuca. In Celosia the inheritance of several colour varieties and of 
fasciation is being studied. In Lathyrus the investigations include a cytological 
study of the pollen development, with particular reference to (a) the dominance 
of long pollen; (b) the inheritance of pollen sterility; (c) the cytological basis 
of crossing over. With wild cabbages a study is being made of the variability, 
the effects of cultivation, and the self-sterility. In Viarella cordifolia, a form 
with orange anthers is found to be usually almost wholly sterile in its pollen, and 
the ovules are also often imperfectly formed. Further studies of this condition 
are being made. Numerous species of Lactuca are being grown for a com- 
parative study of their chromosomes and external characters. 
In the Oenotheras, which probably show more genetic and _ cytological 
peculiarities than any other groups of organisms, a number of problems have 
reached definite conclusions which link up their behaviour in various ways with 
the more usual Mendelian inheritance. In a paper on present problems of 
Oenothera research which is now in the press in the Mendei Memorial Volume of 
the Prague Celebration, it is pointed out that the differences between the twin 
hybrids frequently obtained in crosses are probably all determined by a single 
pair of chromosomes. Much of the differentiation of the group appears to have 
been concerned with this pair. The well-known gametic and zygotic sterility 
of this group can also be accounted for, as well as the persistent heterozygous 
condition of the species, by the presence of a pair of zygotic lethal factors and 
a pair of gametic lethal factors in different pairs of chromosomes. This obviates 
the necessity for assuming that crossing-over occurs, a hypothesis for which 
there is no cytological basis in Oenothera, although there is evidence of such a 
process in Lactuca. The usual presence of 50 per cent. of empty seeds and 
functionless pollen and megaspores can thus be explained by the presence of 
lethal factors arising, as in Drosophila, through mutations. The presence of 
