of Abraxas grossulariata 229 
was badly developed; (2) the internal genital organs were, how- 
ever, ovaries, and contained a few quite well developed eggs. ° 
In consequence of the discovery of these gynandromorphs, one, 
if not both, of them associated with an apparent failure of the 
normal sex-limited transmission, it occurred to me that possibly 
the two exceptional females of family 12.25 described in Journ. 
of Genet. Iv, 1914, p. 15, might also have been to some extent 
gynandromorphic. Both paired with males, and one laid eggs, 
but these proved infertile. I therefore sent the specimens to 
Mr Pierce, and he reported as follows. The specimen which laid 
no eggs had distinctly asymmetrical genitalia, the right side being 
on the whole better developed, while the left, although female in 
character, had all the organs more or less malformed. In the 
specimen which laid about 20 infertile eggs, the external geni- 
talia were asymmetrical and imperfect, and Mr Pierce is of 
opinion that certain structures may perhaps represent rudiments 
of male organs. 
These facts place in a somewhat new light the conclusions 
with regard to exceptions to the normal sex-limited inheritance 
arrived at in the paper referred to (Journ. of Genet. Iv, 1914). 
Tt was there concluded that failure of the normal sex-limited trans- 
mission of characters might involve sterility, and support was given 
to this idea from the frequent sterility of the tortoise-shell tom-cat 
(Proc. Camb. Phil. Soc. Xvul, p. 8307 and Journ. of Genet. v, 1915, 
p- 65). It now appears, however, that in two cases of apparent 
failure of sex-limited transmission (if the specimen in family 14.31, 
which contained ovaries, is so regarded), the individual is in reality 
a gynandromorph, and that the sex-limited characters are normal 
as regards one part of the sex of the specimen. And further, in 
both the cases previously described as exceptions to sex-limited 
transmission (family 12.25), although the whole insect was pre- 
dominantly fernale, yet on one side the external genitalia were 
abnormal, and in one case showed indications of male structure. 
It is possible, therefore, that both these insects were to a very 
slight extent gynandromorphice, and that the grosswlariata pattern, 
where lacticolor was to be expected, is connected with the pos- 
session of a certain amount of male tendency in individuals which 
were preponderatingly female. If such a view be adopted, it 
would mean that hitherto no true exceptions to normal sex-limited 
inheritance are known in Abraxas grossulariata. 
