_ WILLARD: CRANIAL NERVES OF ANOLIS CAROLINENSIS. 3a 
as the m. depressor palpebrae inferioris. Bradley (:03), in analyzing 
the musculature with a view to explaining its relation to the mastica- 
_tory movements, recognizes in several of the lizards a double function 
for this muscle, as suggested by the fact that some of its fibers are 
inserted on the lower eyelid, while others appear to have their inser- 
tion in the “fibrous membrane circumscribed by the pterygoid, pala- 
tine and the transverse bones”’ (p. 481). This is also brought out more 
fully by Bruner (:07),! who, in looking in the head musculature for an 
apparatus to control the flooding and distension of the orbital blood 
sinuses, discovered that a part of this muscle was completely differ- 
entiated into a m. protrusor oculi, and records its occurrence in eleven 
lizards, including Anolis. It is apparent, however, from a study of 
several series of sections that the two are not completely distinct 
morphologically in Anolis, as the following description will indicate. 
It seems probable, however, that the two functions as described by 
Bruner are here subserved. The following is the condition in Anolis: — 
the thin layer of striated muscle (Plate 3, fig. 7; Plates 4, 5, figs. 
11-15, protru. oc., dep. palb. if.) which lines the floor of the orbit 
ventromesad to all other orbital structures has two distinct origins; 
_ one (protru. oc.) is from a slender tendon attached to the membranous 
| wall of the cranium just anterior to the bony part on a level with the 
| Gasserian ganglion (Plate 3, fig. 7; Plate 6, fig. 16). This tendon 
| passes down mesad to the columella (epipterygoid) and is continuous 
| with a muscle band which lies closely applied to the membranous 
|region (Plate 5, figs. 14-15) referred to by Bradley. While some of its 
| fibers may be inserted here, most of them continue and spread out to 
form the anterior part of the broad palpebral muscle with insertion 
(on the lower lid. This is best shown in the dissection (Plate 3, fig. 7). 
'The second origin, ventral to that of the first, is from the fascia on the 
ventral face of the bursalis muscle; this band crosses ventral to the 
posterior rectus muscle (Plate 8, fig. 7; Plate 5, fig. 15, dep. palp. if.) 
and turns sharply to form the posterior half, or more, of the palpebral 
fnusele. The two bundles of origin include between them the jugular 
vein and suborbital sinus of the same side. The innervation of these 
jauscles favors the view of a double function, and suggests for a part 
; f it at least a more visceral function, such as that assigned to it by 
“pruner. This will be described in detail in connection with the ac- 
_ount of the nerves (p. 50). 
| 
' Bruner does not mention the earlier recognition of this muscle by Bradley. 
