46 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 
tively simple. The ganglion (Plate 5, fig. 15, gn. cil.) lies between the 
membranous cranium on one side and the bursalis and retractor oculi 
muscles on the other, and is separated from both III and V by roots _ 
of considerable size. It is connected with nerve III, as is generally 
the case in other forms, by a shorter, thicker root, radix brevis (Plate 2, 
fig. 4, rx. cil. II), while the root from V (rz. cil. V) is longer and more 
slender. Only the proximal end of this root is shown in the figure. 
Both roots communicate with the ganglion directly and in like manner 
the two ciliary nerves arise directly from its distal end. .In the 
plotting (Plate 2, fig. 4) this point is hidden by nerve III. These 
ciliary nerves (Plate 5, figs. 13, 14, cil.) cross dorsad of the optic nerye 
to enter the sclerotic coat of the eyeball. They keep close together 
in their course as they pass laterad and cephalad around the eyeball 
to their terminations in the striated muscles of the ciliary body and 
epithelial surfaces. From the smaller of these ciliary nerves is given 
off a very fine branch, which follows the others for a distance but is 
lost before its entrance into the eyeball. In sections individual 
medullated fibers are seen to be given off from this small ramus. 
These are lost along the walls of the small blood vessels. The pres- 
ence of these fine rami in immediate contact with the retractor oculi 
muscle at its end of insertion could readily give rise to a misinter- 
pretation in regard to their distribution (see p. 32). It is possible 
that Osawa (’98, p. 537), basing his conclusions on dissections alone, 
made such an error in his account of Hatteria. In Anolis every 
recognized fiber leaving the ciliary nerves could be traced cephalad 
beyond the most anterior extent of this muscle (tr. oc.), and in no case 
were these fibers distributed to the muscle. | 
In many points the microscopic evidence was tu from conclusive — 
for determining the relations of the ciliary roots and nerves to the 
ganglion, but they offer certain facts worth recording. The short — 
root is a large one and is principally composed of fibers of very light — 
medullation, but not of the smallest caliber, 7. ¢., they are larger than 
those of the visceral sensory system as shown in palatine VII. Among 
these are a very few coarser fibers of a medullation sufficiently heavier 
to make them conspicuous. These are as large as many that remain 
in the somatic motor rami, but do not equal in size those which pre- | 
dominate in these motor nerves, nor are they segregated into a single » 
group to be readily followed through the ganglion. However, the 
fact that every section through the ganglion shows about the same | 
number of these coarser fibers points to the possibility of their unin-— 
terrupted passage. The ganglion is a uniformly oval structure, the — 
cells of which entirely surround the short root. 
