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WILLARD: CRANIAL NERVES OF ANOLIS CAROLINENSIS. 47 
The longroot (rx. cil. V.), from the nasalis division of the opthalmic, 
is composed of both medullated and non-medullated fibers and joins 
the ganglion on its dorsal side (Plate 5, fig. 15) about midway of its 
length, many of its fibers immediately mingling with the ganglion 
cells. The independence of the bundle is, however, preserved through- 
out by the continuity of the non-medullated components of this root, 
which passes across the dorsal side of the anterior half of the ganglion 
and on into the large ciliary nerve, in which it appears for a\con- - 
siderable distance as a lighter area in the cross section. These facts 
do not preclude the possibility that many of the non-medullated 
fibers end in the ganglion, nor that a part of the postganglionic bundle 
of fine fibers takes origin in the ganglion. Further analysis of the 
ciliary nerves discloses some of the coarse fibers of the short root in 
each. The rest are of the smaller medullated variety and appear 
to take their rise as peripheral neuraxons of the cells of the ciliary 
ganglion, since they do not occur in the ciliary roots. 
In comparing Anolis with other Sauropsida we find that the arrange- 
ment of roots, ganglion and ciliary nerves is that which Fischer (’52, 
p. 117) describes as typical for reptiles. As an exception he mentions 
Salvator merianae, where the trigeminal and oculomotor roots join 
proximal to the ganglion. Other accounts agree with Anolis. Osawa 
(98, p. 602), in describing Hatteria, establishes another exception, 
wherein the ganglion itself is not connected with the trigeminal nerve 
by an independent root, but, if at all, through recurrent fibers, as in 
jthe fowl, the ciliary rami from V joining the ciliary nerve distal to 
ithe ganglion. 
Carpenter’s (:06, p. 158) careful analysis of the ciliary ganglion 
and its connections in the adult fowl is the only basis we have for a 
somparison of histological features with conditions in birds. In the 
‘owl, the short root from the oculomotor being much the same as for 
\nolis, one main ciliary nerve leaves this ganglion. This contains all 
he well medullated fibers. Another small bundle, of feebly medul- 
ated fibers, leaves the ganglion dorsal to the large one. A third 
small) ramus accompanies the other two; microscopic study, how- 
_|ver, showed Carpenter that it contains no fibers from the ganglion, 
ut is merely a communicating ramus from the trigeminal, which 
sets the ciliary nerve distal to the ganglion. The same ramus 
nds some recurrent fibers back to the ganglion. Other fine rami 
a be given off from the communicating ramus. All the trigeminal 
ements are non-medullated. The ciliary ganglion itself is divisible 
to a sympathetic and a cerebro-spinal part. 
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