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WILLARD: CRANIAL NERVES OF ANOLIS CAROLINENSIS. dl 
by any writer to a muscle innervation similar to the one above de- 
scribed. This peculiarity, coupled with the described function (p, 
33) of this muscle, offers a problem for comparative study. It is 
to be noted that no ganglion cells were found at the point of union 
either on V or VII. 
Afferent neurons. ‘The central course of these fibers is so involved 
in other brain tracts that the present description will begin at the 
superficial origin of the root. The descending root of the trigeminal 
nerve traced cephalad approaches the surface of the medulla and 
produces a swelling of the surface beneath the roots of nerves VII and 
VIII (Plate 6, fig. 18, rx. V). The motor neurons, as before described, 
join this bundle rather abruptly from a more median position, and the 
two together leave the brain as a single root. The fibers making up 
the sensory components of the trigeminal lack uniformity of size and 
medullation. They are, however, of a size approximately that of the 
motor components of the trigeminal and facial, but also have among 
them both larger and smaller fibers. A characteristic feature of the 
cross sections is the presence of a few very large, strongly medullated 
fibers. These are recognizable in the brain, and a central connection 
may possibly be established for them. Peripherally they seem to 
offer no special relation to end organs. They occur in limited num- 
bers and are always scattered as isolated fibers. It is possible they 
are motor in their character and are transferred to the motor rami. 
The ophthalmic, maxillary, and mandibular divisions of the trigemi- 
nal are recognizable even proximal to the ganglion (gn. V; gn. opth.), 
the two ganglia being entirely distinct and scarcely in contact in 
series 30. The relative sizes and positions of the ganglia are best seen 
from the figures on Plates 2, 3. The cross section (Plate 6, fig. 17) 
shows the segregation of the two cell groups. In addition to these 
two ganglia, one of the figures (Fig. 6) shows a third; a very small ven- 
tral group of cells entirely similar in the sections to those of the larger 
ganglia. Careful study was made of this small ganglion to discover 
any structural features that would indicate a sympathetic character, | 
but comparison showed no such group on the opposite side of the 
same individual; in this case therefore it is probably a separation (in- 
constant) of a few cells of the main ganglion. From what we know 
_ of other forms, it seems probable that the sympathetic rami of the 
lachrymal plexus send fibers to the Gasserian ganglion, but that could 
not be demonstrated in Anolis, and there is no deep sympathetic trunk 
connecting the Gasserian with more posterior ganglia. 
A. detailed account of the peripheral distribution of the sensory 
