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WILLARD: CRANIAL NERVES OF ANOLIS CAROLINENSIS. 53 
ramus. The frontal ramus lies dorsal to the other and both run 
cephalad for some distance, passing between the membranous crani- 
um and the ligamentous origin of the protrusor oculi muscle (Plate 6, 
fig. 16,f.andna.). A little farther forward the frontalis turns abruptly 
dorsad to the posterior angle of the orbit (Plate 5, fig. 15). In this 
course it keeps next the membranous cranium and attains a position 
just caudad to the lachrymal gland. As it turns dorsally it gives off a 
branch that becomes involved in the sympathetic plexus (Plate 3, 
figs. 6, 7), but, like the other branches, this contains cutaneous sen- 
sory elements that can be traced to the skin. The main branch con- 
tinues between the brain and the large blood sinus which lies just 
posterior to the lachrymal gland (Plate 5, fig. 15). A little farther 
forward, and at a level where the gland is beginning to be cut, there 
are given off a number of branches, which anastomose with the 
sympathetic, but for the most part supply the upper lid. These lie 
just mesad of the lachrymal gland (Plate 5, fig. 14, gl. Ich.). The main 
part continues forward and supplies, from time to time, small rami 
to the skin between the eyes (Plate 5, figs. 12, 13). 
Ib. Ramus nasalis (na.). This is the main ophthalmic branch and 
takes the characteristic course through the orbit (Plates 4, 5, 6, figs. 
10-17). It passes into the orbit with nerve III and passes dorsally 
to the optic nerve and over the ocular face of the dorsal rectus mus- 
cle (Plate 5, fig. 14). Just before its entrance into the orbit (Plates 
2, 3, figs. 4, 6, rx. cil. V.) it gives off the long root of the ciliary nerve 
(see p. 46). With this exception there are no branches or connec- ° 
tions within the orbit. It passes out of the orbit on the median side 
of the Harderian gland (Plate 3, fig. 7; Plate 4, fig. 11) into the space 
bounded by the anterior median wing of the palatine, the frontal, 
' the prefrontal and the origin of the ventral oblique muscle (Plate 4, 
fig. 10). Here the nerve passes through the “ ethmoidal’’ sympathe- 
tie ganglion (gn. eth.), which will receive especial description. This 
ganglion lies on a level with the anterior boundary of the bony orbit, 
and through it a cutaneous branch of coarse fibers is given off to the 
| skin of the anterior angle of the eyelids (Plate 4, fig. 10, na.!). Soon 
after this the main nerve divides into a lateral branch (Plate 4, fig. 9, na. 
| L) and a median branch (na. m.). These richly supply the skin of the 
snout. The median one gives off in addition a branch to the mucous 
/membrane of the nose; its cutaneous branch passes forward along 
7. —_ line to be distributed to the end of the snout (Plate 2, 
| ig. 4). | 
| According to Watkinson (:06, p. 458), who mentions both the fron- 
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