WILLARD: CRANIAL NERVES OF ANOLIS CAROLINENSIS. 5S 
An anastomosis of palatine VII with the ophthalmic branch of V in 
the anterior orbital region seems to be of wide occurrence in the Saurop- 
sida and Amphibia. It is described in Amblystoma (Herrick, ’94), 
tadpole of the frog (Strong, ’95 [farther forward here]), Spelerpes 
= = a 
———— 
(Bowers, :00), Amblystoma (Coghill, :02) and Amphiuma (Norris, 
108). “ 
Of the above mentioned investigators, Coghill finds a ganglion at 
the point of union, and Norris (p. 540) says “there seems to be a 
ganglion on the palatine nerve shortly before the anastomosis”; be- 
yond the anastomosis he finds mixed cutaneous and viscero-sensory 
rami. With this observation by Norris agrees the statement I have 
made regarding Anolis; that this ganglion, while surrounding ophthal- 
mic V, is really more closely related to palatine VII. 
Among the reptiles an anastomosis is mentioned by Gaupp (’88), 
Osawa (’98), and Watkinson (:06). .On the other hand Fischer (’52) 
does not mention it at all. The presence also of a ganglion, at least 
one discernible by dissection, seems to be less general in the described 
reptiles than the condition in Anolis would suggest. The only men- 
tion of it which I have found was in Gaupp’s (’88, p. 469) description 
of the snake Tropidonotus, in which he refers to it as “eine leichte 
Anschwellung,” and further “Vielleicht entspricht dieselbe dem 
ganglion ethmoidale der Végel.”” Watkinson (:06, p. 463) speaks of 
palatine VII and nasalis V as lying quite close together at a point 
corresponding to the ethmoidal ganglion and having one or two con- 
necting branches. It is probable that sections would have disclosed a 
ganglion here. She found no cutaneous branch of the nasalis at this 
point. 
Peripheral distribution of the ramus nasalis. The distribution of the 
cutaneous sensory fibers carried in this nerve begins with the bundle 
of coarse fibers entering the posterior end of the ethmoidal ganglion. 
Distal to the ganglion the main nerve divides into the lateral and 
median branches before mentioned (Plates 2, 3, figs. 4, 6, na. L., 
and na. m.), which are of about equal size. One of these retains a 
median position while the other crosses the roof of the nasal capsule 
to a more lateral one. 
(a) Ramus lateralis nasi. This ramus in passing cephalad and 
laterad across the roof of the olfactory chamber occupies a position 
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between the cartilaginous capsule of the chamber and the cranial 
bones. It is distributed to the skin over the subterminal region of the 
_ snout, that is, just posterior to the external nares. Before reaching 
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the integument the lateral ramus passes through the nasal gland, 
