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68 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 
connection between the superficial sympathetic trunk of the head and 
the infraorbital nerve. 
(2) Ramus hyomandibularis (hy-md.). This nerve leaves the genicu- 
late ganglion (Plates 2, 3, figs. 4, 6, 7, gn. VII) at its posterior end. 
It embraces viscero-motor and viscero-sensory components. In eross 
sections it is to be seen that these two components are entirely sepa- 
rate throughout their course, the motor fibers having the more dorsal 
position. The latter include all the motor elements of nerve VII; 
these pass from the root across the dorsal side of the geniculate gan- 
glion, not penetrating it, and then turn sharply into the ramus hyo- 
mandibularis. ‘The motor and sensory elements each take up about 
half the area of a cross section of the nerve, the sensory components 
being of course much finer in caliber than the others. The course of 
this ramus is dorsad and caudad, following the same projecting ledge 
of the prootic bone (Plate 1) as does the palatine, but in an opposite 
direction (Plate 3, fig. 7). This course leads it to a point just ventral 
to the articulation of the quadrate with the prodtie process of the 
ear capsule (Plate 6, 7, figs. 19-20). Here occur the crossing and 
anastomoses of the head sympathetic trunk (comn. ex.) with the hyo- 
mandibular ramus of VII (Plate 3, fig. 7). The superficial sympathetie 
trunk (comn. ex.) from this point to the lachrymal plexus and infra- 
orbital ganglion is called by Watkinson (:06, p. 464) “ramus recurrens 
nervi trigemini ad facialem”’; its continuation to nerve IX, the “ramus 
communicans externus cum glossopharyngeo.”’ 
In a series of sections of Anolis, in which fixation in this region was 
excellent, it is shown with certainty that practically all the medul- 
lated fibers in this sympathetic ramus, which in every way resemble 
viscero-sensory fibers, pass the facial nerve in continuous course. 
The facial, however, makes its way between the sympathetic fibers, 
most of which cross the facial nerve mesally; a few only split away 
from the others to cross it laterally, and then immediately rejoin the 
main bundle. This is not a sympathetic center of any importance, 
although from four to six very small ganglion cells are inclosed between 
the two nerves at the point of crossing. While there was no evidence 
that any of the fibers in the part of the sympathetic ramus between 
VII and V turned into any of the parts of VII at this point of union, 
there does seem to be a strong indication that some of the fine medul- 
lated fibers contained in the hyomandibular ramus of VII are continu 
ous with a part of those in the posterior section of this sympathetie 
trunk, i. e., that between VII and IX. 
Attention is here called to the fact that cutaneous fibers are not 
