Dr. C. W. Andrews — Neio Eocene Crocodiles from Egypt. 483 



Crocodilus sp. 



From the Middle Eocene beds also remains of a broad-snouted 

 crocodile have been collected, but they are so fragmentary that 

 further material is necessary before any attempt to distinguish the 

 species is made. 



Of the genus Tomistoma one species, T. africanum, from the Middle 

 Eocene (Qasr-el-Sagha Beds) of the Fayum (Geol. Mag. [4], 

 Vol. VIII (1901), p. 443), has already been described. Now two 

 others, one of slightly later, the other of rather earlier date, are to 

 be considered. 



Tomistoma. gavialoides, n.sp. 



In the Upper Eocene beds of the district remains of a gavial-like 

 crocodile are fairly common. One nearly complete skull has been 

 collected by Mr. Beadnell, and it is to this that the following 

 description refers. This species is interesting because in some 

 respects it is intermediate between Tomistoma and Gavialis, though 

 much nearer the former, its relationship with which is especially 

 marked in the comparative fewness of the teeth (though they are 

 more numerous than in T. Schlegeli) and the extension forwards of 

 the nasals to the premaxillee, between the facial processes of which 

 they run for some distance. 



The form of the occipital surface both above and below the 

 foramen magnum is almost exactly as in Gavialis, and, as in that 

 genus, the skull roof is wider than in Tomistoma Schlegeli, owing 

 mainly to the large size of the supra-temporal fossas. The orbits 

 differ from those of the recent Tomistoma in being more rounded, 

 as in Gavialis, but at the same time they have not the prominent 

 borders found in that genus ; in the breadth of the inter-orbital bar 

 also the skull is exactly intermediate. The snout narrows less 

 suddenly than in Gavialis, but not quite so gradually as in Tomistoma 

 Schlegeli; as in this latter the nasals extend forward to the pre- 

 maxillse, in Gavialis they are short and separated from the pre- 

 maxillee by a long interval. The expansion of the premaxillary 

 region is slight, less than in the true Gavials. 



There are 22-23 teeth on either side ; in T. Schlegeli the number 

 is 20-21, in Gavialis 27-29. The premaxillse bear five teeth, as in 

 the Gavial ; in the recent Tomistoma there are only four, though in 

 some fossil species referred to this genus there are five. The 

 maxillary teeth seem to have been almost equal in size throughout 

 the series, and to have been directed somewhat outwards and forwards, 

 both Gavial-like characters. 



This skull resembles in many respects that described by Toula 

 and KaiP under the name Gavialosuchus eggenhurgensis. This 

 species also is in many ways intermediate between Tomistoma and 

 Gavialis, but has been referred by Lydekker to the former genus. 

 It differs from the present species in the narrower skull roof, the 

 smaller temporal fossae, and the raised borders to the orbits ; but at 



1 * ' Uebef einen Krokodil-schadel aus den Tertiarablagerungen von Eggenburg"; 

 Denksch. Akad. Wiss. AVieu, Bd. l (1885), p. 299, pis. i-iii. 



