84 MR. R. F. TOMES ON A NEW BAT. [Feb. 24, 



05s.— Since the above was written, I have made a careful compa- 

 rison of the skull of this singular Bat with that of several other 

 species hitherto doubtfully placed with the Phyllostomidae. The 

 following are the results:— The genus Schizostoma, which is rather 

 intimately allied to Vatnpyrus, bears also considerable resemblance 

 to the genus Macrotis in the general conformation of the cranium 

 and the lower jaw, and also in the very great similarity in the den- 

 tition. The form and size of the ears, too, in these genera are very 

 similar. Macrotis, again, bears in several particulars an intimate 

 relationship with the present genus Lonchorhina. More especially 

 may be mentioned the length of the tail, which extends in both ge- 

 nera to the whole length of the interfemoral membrane, as in the 

 genus Vespertilio, the considerable development of the ears, the size 

 and freedom of the feet, and, perhaps more than all, the general 

 contour of the cranium. 



Pursuing the comparison, we find that Lonchorhina bears very 

 considerable resemblance to Chilonycteris, in the form of the anterior 

 part of the cranium, in the number and relative size of the teeth of 

 both jaws, and in the form of the lower jaw. Passing on from Chi- 

 lonycteris to Mormops, the skull of the latter is seen to be an exag- 

 geration of the former, having the facial part still more depressed, 

 and the cerebral part still more elevated. The upper teeth in both 

 these genera are very similar ; and those of the lower jaw do not 

 present any essential differences, the chief one being that in Chilo- 

 nycteris the middle premolar is very much smaller than the corre- 

 sponding one in Mormops, which, although smaller than those on 

 either side of it, is not minute. All the above-mentioned genera 

 agree with each other in the presence of a fourth joint to the longest 

 digit of the wing, and in fact must be said to bear considerable re- 

 semblance in most particulars, saving in the degree of development 

 of the tail and the existence or absence of a hastate nose-leaf. How- 

 ever, it may be said that those species which have not a nose-leaf re- 

 sembling that of the ordinary Phyllostomidae have nevertheless some 

 cutaneous development about the face, nose, or mouth, and cannot 

 be properly called simple-nosed species. 



There is another very singular genus, of which I have before spoken 

 in commvmications to the Society, and which I have regarded as 

 allied to Molossus, but I have mentioned that it possesses four pha- 

 langes in the longest finger. I allude to the genus Mystacina, which 

 has\itherto been found only in New Zealand. When preparing my 

 paper on the Bats of that country, I had not examined either Mor- 

 mops or Chilonycteris ; but, on afterwards working out some West 

 Indian Bats, was at once struck with certain resemblances between 

 the latter and Mystacina. Without at present alluding to the de- 

 tails of structure which have induced me to arrive at this conclusion, 

 I take this opportunity of stating that I now regard Mystacina as 

 an aberrant form of Phyllostomidae, coming after the several genera 

 which have been compared with each as above, but differing more 

 from them than they do from each other. 



