Dr. Henry Woodward — On a new Loricula. 499 



much more specially developed than the others, and the hard external 

 covering was more completely divided into two series ; the upper 

 series being more developed to form the plates of the capitulum, 

 whilst the remaining rows, nearly all alike in form, constituted the 

 scales of the peduncle. It is here that we have observed a modifica- 

 tion in a form belonging to the Cenomanian [Loricula Syriaea, Dames) 

 and in another form from the Upper Chalk {^Loricula pulchella, Sow.). 

 The evolution beginning in this way becomes more and more apparent, 

 in such a manner as to constitute finally the forms with which we are 

 familiar in both a fossil and recent state, as Pollicipes for example, 

 in which two verj' distinct general regions exist, one above forming 

 the capitulari-an plates, which are well developed and most thoroughly 

 protect the soft parts of the animal, and another lower region, serving 

 solely as a support and covered by the peduncular scales, nearly all 

 alike and much smaller in size than those of the capitulum. 



" The above deductions are supported clearly, not only by 

 palseontological proofs which we shall specify, but also facts of 

 embryology and anatomical details. 



' ' Therefore, if, for example, one studies the post-larval development 

 of Pollicipes polymerus, Sow., one sees that in very young specimens 

 it is impossible to define the capitulary region because the lower 

 plates of the capitulum nearly resemble the upper scales of the peduncle. 



"Again, Koehler was the first to point out an organ in the centre 

 of the scales of Pollicipes, which we have described after him as 

 being an organ of sensation. One finds this again, modified it is true, 

 quite in the inferior plates. It only disappears in those in which the 

 scales have arrived at a much higher stage of development. 



" We see, on studying each of the sections of this sub- class, how 

 the different forms are derived from the primitive type and how they 

 are connected again one with another." (Gruvel, Introduction, pp. 5, 6.) 



This large Loricula, both on account of its much greater size and 

 more remarkable capitulum, merits recognition as a new species, rather 

 than to be placed with Loricula pulchella or only treated as a variety. 

 If such distinctive points as the form of the scutum and the latera 

 deserve specific recognition,' I desire to name it after the illustrious 

 author of the monograph on the Cirripedia [published by the Ray 

 Society, 1854], and of the fossil Lepadidae and Balanidse [by the 

 Palseontographical Society, 1851 and 1854], Loricula Barwini, — 

 whose great work on "The Origin of Species by Natural Selection" 

 has just been celebrated by the Linnean Society of London. 



I am much indebted to my friend and former colleague for so 

 many years, the present Keeper of the Department of Geology, 

 Dr. Arthur Smith Woodward, F.R.S., for giving me the pleasure of 

 describing this fine, newly discovered Loricula from the English Chalk, 

 which has now been added to the National Collection. 



1 A comparison of Loricula Barwini (Fig. 1, supra, p. 493) with that of 

 Z. pulchella, var. minor (Fig. 2, p. 494) wiU emphasize the specific differences 

 between the former and the latter. Fig. 2 agrees closely with Darwin's original 

 figure of L. pulchella (Mon. Pal. Soe., 1851, T. 5), especially in the outline of the 

 valves of the capitulum, and both differ markedly from the capitulum of L. Larwini, 

 as drawn in our Fig. 1. 



