90 THE MAMMALIA OF THE DEEP KIVER BEDS. 



A fragment of the pes accompanies the type specimen, which, however, displays 

 no features of especial interest, as the hones are not sufficiently well preserved to 

 show the minor changes in the articulations, which are so important in the equine 

 series. As in the American genera of this series, the ento- and mesocuneiforms are 

 coossified, not as in the European species of Ancliitherium, the ecto- and mesocunei- 

 forms. The internal cuneiform is very large, and forming nearly a right angle 

 with the median, extends heneath the entire plantar horder of the navicular to the 

 cuboid. The ectocuneiform is higher vertically, in proportion to its breadth, than in 

 Protoliippus. The jiroximal portion of the median metatarsal is rather slender and 

 rounded ; that of the laterals is surprisingly large antero-posteriorly, but in part, at 

 least, this is due to crushing. The shaft of the laterals rapidly tapers and becomes 

 very slender. The median metatarsal appears to have a slight contact with the meso- 

 cuneiform, but the specimen is too imperfect to determine this point with certainty. 



A second specimen, consisting of the tarsus and portions of the metatarsus, 

 should probably be referred to this genus, though possibly representing a diiferent 

 species. It differs from the fragmentary pes belonging to the type specimen in only 

 one particular, viz., in the much narrower proximal end of mt. ii. Part of this differ- 

 ence is no doubt due to the crushing to which the type specimen has been subjected, 

 but not all of it, and the remainder may be referred to either individual or specific 

 variation. Compared with the tarsus of Protoliippus sejunctus, which Prof. Cope has 

 kindly lent me for the purpose, some not unimportant divergences may be observed. 



In size and general appearance the two specimens closely coincide; the differ- 

 ences are in matters of minute detail and are especially to be found in the relative 

 development of the various facets. On the calcaneum attributed to Desmalippus the 

 additional facet which runs distally from the ectal astragalar facet is somewhat 

 longer than in Protoliippus, but is not so clearly demarcated from the main facet. 

 As in the latter, the ectal astragalar facet is in contact with that on the sustentacu- 

 lum, but this latter surface is considerably broader and more nearly perpendicular to 

 the long axis of the bone. The cuboidal surface is shorter, less distinctly divided 

 into two parts, and at the plantar end less incurved, and is thus separated by a wider 

 interval from the sustentacular facet. 



The astragalus displays corresponding differences. The notch for the ectal cal- 

 caneal facet is wider and its distal continuation longer ; from the latter the inferior 

 border rises abruptly and terminates, as in JEquus, in the beak formed by the sudden 

 termination of the external astragalar condyle. In Protoliippus sejunctus there is no 

 such beak, but the outer condyle curves gently and without interruption into the 

 external plantar border and the accessory calcaneal facet. Whether this is true of 



