44 CHANDLER—DONATIA NOVAE-ZELANDIAE. 
petalous corolla and free stamens, appear to him without doubt 
to have relationship, from the fact that the leaf-structure in the 
two subfamilies agrees, and both have inulin as a reserve material. 
Their general habit also is alike. There is no doubt that except 
for the extrorse anthers the evidence is all in favour of Donatia 
being placed in the alliance of the Campanulateae rather than . 
in the family of the Saxifrageae, as was the case in the older 
classification ; but the justification for Mildbraed’s further step 
of combining the subfamily Donatioideae with the subfamily 
Stylidioideae in the subfamily Stylidiaceae is not at all evident. 
The differences between these two groups seem as great as, if not 
greater than, those between any of the families of the Campanu- 
lateae. Compare, for example, the Campanulateae and the 
Stylidiaceae : in the former we have the flower actinomorphous, 
5-merous, sympetalous, except for the 3-merous gynoeceum, 
the anthers are free, introrse, and there is one style with collecting 
hairs: in the latter the flower is zygomorphous, 5-merous in peri- 
anth, sympetalous, and with two stamens united with the style in 
astylar column. Inrelation to these Donatia shows a flower which 
is actinomorphous, choripetalous, 5-merous in the perianth, the 
stamens 2-3 free from the style, and the ovary divided into 2-3 
chambers. Such evidence might warrant, surely, Donatia being 
regarded as a distinct order, Donatiaceae, of the cohort Campanu- 
lateae, rather than as a subfamily of the Stylidiacea. 
No observations seem to have been recorded of the morphology 
of Donatia novae-zelandiac (Fig. 1), andI therefore take opportunity 
to make known a few facts which I have been able to observe 
in material of the plant obtained from specimens sent to the 
Regius Keeper, Royal Botanic Garden, Edinburgh, by Dr. 
Cockayne, who collected it on the heights of Stuart Island, New 
Zealand. 
Srem.—In a transverse section the stem. of Donatia novae- 
zelandiae shows (Fig. 3) a central mass of xylem with fairly large 
development of cortex and cork. The pith is relatively small, 
and composed of large slightly thickened cells. Surrounding the 
pith is the xylem, composed of regular tracheids—spiral, reticular, 
and annular. There is little xylem parenchyma. Where a 
side branch is given off, the xylem tracheids are elongated in a 
raG@ial direction. Phloem is found in patches around the xylem. 
An endodermis, of sometimes one distinct layer, sometimes more, 
of: oblong thickened cells with granular contents, is present. 
Usually these cells lie adjacent, but often project slightly one 
towards the other (Fig. 3d). The cortex is largely developed. 
The cells composing it are round and fairly uniform in size, 
slightly thickened and loosely arranged with large intercellular 
spaces, No resiniferous secreting system such as is said to occur 
em 
