CHANDLER—DONATIA NOVAE-ZELANDIAE. 45 
in Donatia magellanica* is visible here. Bounding the cortex 
on its outer side are two or three rows of cork cells. They are 
different from those of the cortex, being squarish in shape, with 
thickened walls, and they are filled with oily and granular con- 
tents. From the outermost layer, at intervals, obliquely septate 
hairs are given off, the basal cells of which contain oily and 
granular matter like the cork cells (Figs. 3 and 4). 
Harrs.—The characteristic septate hairs of Donaita novae- 
zclandiae are silky in appearance, and form a dense matting at 
the base of the leaves which are closely adpressed to the stem. 
The ends of these hairs are sharp and bayonet pointed (Fig. 4a). 
The cell forming the point is long. The septa are oblique and 
distinctly pitted and perforated, slightly overlapping on either 
side at the junction of two cells (Fig. 40). The cells on either side 
of such a junction are empty and colourless, but occasionally 
they arethickened. The hairs are comparatively long, and consist 
generally of ten or twelve cells. The basal—one to three—cells 
are cork cells and are quite difierent from the others. They are 
smaller, have horizontal septa, are brown in colour and have a 
distinct granular content, a nucleus, and a thickened wall. The 
lowermost of the basal cells is the shortest, and the others are 
gradually longer until they merge into the empty, colourless 
cells (Fig. 4d). The whole structure of the hair is unique, and 
the presence of the perforated oblique plate-septa may, in a 
xerophytic herb like Donatia, possibly be for the purpose of water 
absorption. Among these hairs is found an epiphytic fungus to 
which I shall refer later. 
Lreaves.—The leaves are packed closely round the stem, 
and are densely covered with the silvery grey hairs at the base, 
which is not narrowed into a petiole. The leaf changes in 
anatomical character from its base to its tip. 
_ At the base, where it is attached to the stem, the leaf has an 
irregular triangular outline, with one vascular bundle in the 
centre (Fig. 6). The epidermis which bounds the leaf is formed 
of small, regular cells with a thick cuticle. A few stomata occur 
on the under surface, and these have no subsidiary cells, differing 
in this also from what is known of Donatia magellanica.— The 
mesophyll is not differentiated, the leaf being symmetrical, and 
consists of slightly-thickened cells, somewhat collapsed to the 
corners of the triangle. Owing to crushing, these cells are some- 
times locally thickened and infolded, particularly at the corners, 
somewhat after the manner, though not to such an extent, as that 
of the infolded mesophyll of Pinus sylvestris. The vascular bundle 
is irregular in outline, with no bounding endodermis. The xylem 
* Solereder, Systematische Anatomie der Dicotyledonen, p. 355. 
? Til. 3332 Sent 
