50 CHANDLER—DEHERAINIA SMARAGDINA. 
and thickly covered with whitish hairs on the dorsal surface. 
Where the lobes of the corolla unite, occur small, ligulate pro- 
jections, the so-called staminodes. These are five in number, 
sub-acuminate, and slightly paler green in colour than the petals. 
(Fig. 2). The five stamens are opposite the petals; at first 
closely adpressed to the style (Fig. 2a; b), latér:they bend out- 
wards, and place themselves against the corolla tube (Fig. 2c, a). 
The filaments are broad, and united at the base around the ovary ; 
the anthers are sub-quadrate in shape and extrorse. The con- 
nective is obtuse, truncated ; the pollen saes are situated at the 
apex of the anther, and a white, mealy mass, consisting of cellular 
fibres, at the base. The ovary is flask-shaped, with a long, slender 
style and discoid stigma. It is unilocular, with numerous ovules. 
borne on a central placenta. 
SHooT.—At the close of the flowering season, Deherainia 
smaragdina begins to put forth numbers of new shoots. These: 
shoots, like the flower buds, arise in the axils of the apical leaves. 
of the rosette. The various stages are shown in the figures. 
(Fig. 3, I-V). The older leaves have been cut off, and their 
positions are indicated by the leaf scars. These new shoots are 
very soft, covered with a profusion of silvery-grey hairs, and a 
number of reddish scale leaves at intervals. In a later stage 
the leaf-buds have unfolded, and formed the same rosette that 
characterises the old shoot. The old stem is woody in character, 
covered with reddish-brown hairs, as indicated, and possessing 
but few scale leaves. As the young stem, therefore, begins to 
take on a woody character, its hairs become darker in colour, 
and the scale-leaves gradually wither away. This can be easily 
seen in (Fig. 3, V), where the scale-leaves of the older shoot are 
relatively small and insignificant to those on a younger stem. 
The internal anatomy of the shoot is interesting. A transverse 
section (Fig. 4) of an old stem shows a large development of 
sclerenchyma outside the bundles. The cells of the pith are 
irregular in size, and are filled with starch. They have thick 
walls and are pitted. The intercellular spaces are small. The 
xylem is transversed by numerous medullary rays which are 
broad, and consist of thin cells elongated in the radial direction. 
A well-marked cambium occurs between the xylem and phloem. 
Outside the phloem are patches of sclerenchyma element rays 
with very small lumen. At the point where the medullary 
emerge the sclerenchymatic cells are larger, and the circular 
laminations are crossed by pits (Fig. 5b). The cortex, consisting 
of thin, regular cells filled with starch, is bounded by the epidermis. 
The cuticle is remarkably thick and corrugated on its surface 
(Fig. 5a); this is more clearly shown in a longitudinal section 
(Figs. 6,7). From the epidermis unbranched hairs are given off. 
