CHANDLER—DEHERAINIA SMARAGDINA. 51 
In an old stem these hairs are very markedly thickened, and 
often swollen at the base (Fig. 6). In a young stem these hairs 
are replaced by numerous, stalked glands, the heads of which 
contain cells very similar to the glands which are found on the 
petals and leaves—though these latter are stalkless. Later on, 
in the development of the shoot, when the cuticle becomes thick 
and corrugated, the glands lose their heads, and become thickened 
to form an ordinary septate hair (Fig. 7). 
A section of the young shoot agrees in appearance—except 
for the absence of the marked sclerenchyma—in all respects with 
a section of an older shoot. 
LEAVES.—The structure of the leaves of the various repre- 
sentative genera of Theophrastaceae—Clavija, Jacquinia, Theo- 
phrastea—has been described already by Votch.* Morerecently he 
describes f also the leaf of Dehevainia smaragdina, the plant under 
consideration. His examination gave the following results :— 
“The epidermal cells are wavy in outline, and the cuticle 
is striated. Hypoderm is absent. The stomata are elliptical 
and do not project beyond the surface. Glands are present, 
with sixteen cells in the head. One-cell-rowed hairs to the length 
of ten cells are plentiful on the under side. The palisade paren- 
chyma are only one-rowed, and consist of short cells. The sub- 
epidermal sclerenchyma are present only in single isolated 
groups on the upper side, but form large bundles on the under side. 
Crystals are present in the lower epidermis, The median nerve 
is simple, surrounded by sclerenchyma, which does not pass 
through it, and the phloem surrounds the xylem on three sides.”’ 
examination of the leaf of Deherainia smaragdina confirms 
these observations generally (Figs. 8 and g). I found, however, 
that very often the stomata do project considerably beyond the 
epidermis (Fig. 8a, 92). The glands are stalkless, and are found 
on both sides of the leaf. The hairs, which are plentiful on the 
under side, are confined to the portion of the mid-rib on the 
upper. 
A transverse section of the petiole, like that of the lamina, 
shows a large amount of sclerenchyma surrounding the median 
vascular system, which gradually splits up into three separate 
strands as the base of the petiole is reached. 
Roots.—The root-structure of the Theophrastaceae has not 
been investigated as has that of the stem and leaf,{ but that these 
organs might present points of interest’ equal to those already 
described, is shown by the structure of the root of Deherainia 
smaragdina. : 
A transverse section (Fig. 10) of a moderately old root shows 
* Votch, Dissert. Erlangen, 1903. + Engler, xxxiii (1904), Bot. Jahrb. 
t Votch, loc, cit, Ge 
