52 CHANDLER—-DEHERAINIA SMARAGDINA 
xylem in the centre arranged in a somewhat spiral fashion from 
the centre. The cambium is succeeded by V-shaped patches 
of phloem, and a many-layered pericycle bounded by the endo- 
dermis, completes the central stele. The cortex is remarkable 
for the size, irregular arrangement, and localised division of its 
cells. The piliferous layer is formed of large cells, which divide 
in all directions. 
The root hairs are remarkable for the thickness of their walls, 
especially at their base. If a section be cut somewhat obliquely 
across the base of one of these hairs, the thickness of the walls 
is at once remarked (Fig. 11a). In the young root, the cortical 
cells are more regular than in the old (Fig. 11). 
FLOWER.—The external appearance of the flower-parts of 
Deheraima smaragdina has already been indicated, but one or 
two additional points may be noted. Typically 5-partite, 
6-partite flowers are not uncommon, and, though themselves 
solitary on their stalks, they are usually developed in pairs in the 
centre of the rosette, one flower much in advance of the other. 
The contrast in colour shade between the sepals and petals is 
well seen in the bud. Not only is the throat of the corolla lined 
with hairs which slope downwards, but it is thickly dotted with 
glands visible to the naked eye, as black spots (Fig. 2a, d). 
These glands are also distributed, though not to such an extent 
onthe upper portion of the petals. Hooker * has described these 
glands as “‘impressed dots on the upper surface.”” Otherwise 
they are unmentioned. 
The internal structure of the flower has not been hitherto 
described, but offers many points of interest :— 
The sepals, though simple in structure, show a well-marked 
epidermis on both sides, exhibiting a corrugated, cuticular layer 
on the outer wall. The cells composing the mesophyll are large 
and homogeneous ; the vascular bundles are numerous, and show 
distinct xylem and phloem. Glands are present, are on the 
interior surface, but not in any great number; these sepaline 
glands, in contradistinction to those at the base of the petals, are 
all sessile and sunk to the level of the epidermis (Fig. 12a). 
The petals on the other hand, in cross-section, show the glands 
more thickly distributed than on the sepals. They occur in all 
stages, and their development can easily be traced. They origin- 
ate from an epidermal cell, which divides transversely; the 
upper cell thus formed becomes the head, and the lower, which 
takes no further part in the development, remains unchanged. 
By a series of divisions the upper cell ultimately contains about 
sixteen cells situated in a radiate manner like the spokes of a 
~ wheel. ss | 
a a * Bot. Mag. 1878, 
