- 
1882.] PROF. ST.-GEORGE MIVART ON THE ELUROIDEA. 519 
SrxvuAL ORGANS. 
The penis presents every variety as to its internal osseous support. 
It is entirely boneless in the Hyenide; and the bone is rudimen- 
tary or absent in the Viverrine. It is tolerably developed in the 
Herpestine, and very large indeed in Cryptoprocta. 
Cuvier says' of the Ichneumon :—“ Le gland est comprimé sur 
les cotés, arqué en dessus 4 son extrémité, et composé, en trés grande 
partie, de los qu’il contient. Son bord inférieur présent une fente 
qui ne s’étend pas jusqu’A l’extrémité. Elle aboutit 4 une sorte de 
cul-de-sac trés profond, qui remplit l’échancrure de l’os, et au fond 
duquel viennent s’ouvrir, par deux orifices séparés, l’urétre et le canal 
excréteur commun des glandes de Cowper.” 
Crocuta differs from all other Hluroids in that the clitoris is enor- 
mous and traversed by the urogenital canal *. 
In Aretictis the Cowper’s. glands are each less than half the size 
of the bilobed prostate. In Prionodon, on the contrary, each Cow- 
per’s gland is about as large as is the whole of the bilobed prostate. 
According to Professor Owen (Anat. of Vertebrates, vol. iii. 
p. 780), Felines have usually six nipples, four ventral and two pec- 
toral ; but there are eight in the Domestic Cat. There are four ven- 
tral teats in Paradoxurus, Herpestes, and Hyena; but in Viverra 
there may be two pectoral teats also. 
Scent-GLanps. 
Of these glands there are two very distinct categories. One cate- 
gory comprises the prescrotal scent-glands, and the other the anal 
glands. 
The Prescrotal Glands *, 
These are met with in their most complete form in Viverra civetta, 
where there is a pair of medianly adjoined glands placed between the 
penis and the scrotum in the male, and in an analogous position in 
the female. Their secretion escapes into a large scent-pouch, the ex- 
ternal aperture of which appears as an antero-posteriorly extended slit 
simulating a large vaginal aperture. This aperture is larger than 
either the vulva or the anus. Each scent-gland is about twice the 
size of one of the animal’s anal glands ; and in each scent-gland is a 
central cavity which communicates with the median scent-pouch, 
into which the glands also open by a number of minute orifices. 
In Viverra zibetha* the scrotum is similar. In Aretictis the 
secretion of the prescrotal gland exudes into a naked cutaneous inva- 
gination placed, like a vulva, in front of the anus. A similar inva- 
gination exists in Paradovurus, and a naked space in Nandinia. 
In Genetta and Hemigalea the conditions are those already de- 
scribed and figured by me®. The scent-glands contain numerous 
' Legons d’Anat. Oomp. vol. viii. p. 222. 
2 PZ. 8. 1877, p. 369, fig. 41. 
° Ohatin, Ann. des Sci. Nat. (5th series) vol. xix. (1874). 
* There is an old but an excellent preparation of these glands in the Museum 
of the Royal College of Surgeons, Physiological Series, No. 2514. 
° P.Z.8. 1882, p. 156. 
Proc. Zoou. Soc.—1882, No. XXXV. 35 
