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Part IIl.—On the Morphology and Phylogeny of the Paleognathe (Ratite and Crypturi) 
and Neognathe (Carinate). By W. P. Pycrart. 
Tasty or Conrents. 
Pterylosis Page Nervous System and Sensory Organs. Page 
CHAIN cogossapdolboedsugspQeouboodD 151 IyeriNen ono CODCOD UOGOdBUeCOnOb TauOC . 247 
» Embryo and Nestling ................ 159 YOY ravers atareucter crete oreseolcts eesti Nexeiov seri eis 247 
Structure of Feathers ................-- 162 Jacobson’s cartilages...........0.0-- 000s 247 
WMheENeossopbilesi rare <iaeepelels slate) eleet= 162 Alimentary Canal and Appendages. 
Adult and Nestling Remiges of Casuarius . 165 DBiccalicavitivg | pests syohetecrersrs spel sana ote tete tee 248 
Rel eopttles etspatetererey=letetstoicielsic(oisverer ss-tetere 166 Intestinal convolutions ...............- 248 
Summary of Pterylological Characters .... 168 Liver and Gall-bladder ................ 251 
Respiratory Organs. 
Osteology. ATSTATE OS a yotoneveWrefayavateverctale oianeny state(arcter Yate sie 252 
‘Seull GeUAG bt Te cae copanenonocudacesodc 171 IATTRACH elas de ote ee oe taleisie vistas eee e Mele ote 252 
4 SEIS Spoons ro oapaoboa er 195 Windpipes stijj5cteeiew oct seston ae 254 
Some points of significance in the Struthious Choanse a erect aiid cieiaey cer eroiererstaiaete 256 
HAL UCI. cag omadacno come coo DO coe no 206 Circulatory System. 
VEDA C ITNT 6 dno Sobran RS bOOpICE 208 HV Garth ols, oxeyayoieunja ters aatevorsyens vote elstsiateeeyavainie 256 
UAT et oa Gdu Ute On,ID UOSURODCOOeOOn Sp 215 Caxotldsiy cer rcier-ieriere tetera eta ee 257 
Sternum and Pectoral Arch.............. 216 Cloaca, Bursa Fabricii, and Urino - genital 
LAR oA eatin ntigobncddbangeoes ac 225 System. 
PEGOTAL MSU) tour oteteletelele al-(cletsietelsteleter sts 233 Cloacan ‘cc, <ic-a1e yoversysin oe orate netate tereeo eye 257 
Lai banhinn seagouvoonpeduanunedoodars 235 Bnraauyeatsssenesasedaieles: eid eee cone 258 
Copulatory, Org ari s.. cele ntalcistnieleiarie trate 259 
Myology. DEVELOPMENT o,<) 5 oe oles clalele © «lle ole theta teens 259 
Skin-muscles ..... GORICOCODC OMmome anc 238 Phylogeny of the Paleognathe .............. 259 
Muscles of Wing and Shoulder-girdle .... 238 RU HLO BNO ODO FO Op HEA OE OE t4.0,0.00 ao Oe 266 
Thigh- and Leg-muscles .....0........+ 242 Keys to Genera and Species ..... 00.00.00 es 267 
Inrropuctory. 
Tue present investigation was undertaken at the request and with the co-operation 
of the Hon. Walter Rothschild, M.P., who has furthermore furnished the bulk of the 
material, and in numerous other ways greatly aided my work. The collection of 
skeletons in the British Museum of Natural History has proved a source of great 
wealth. I am besides indebted to Mr. Beddard, Dr. Gadow, Dr. Forsyth Major, and 
Prof. Howes for the loan of specimens in their possession or charge, and I take this 
opportunity of recording my thanks. 
The object of this investigation has been to ascertain, as far so possible, the rela- 
tions of the Casuariide to the remaining “ Struthious ” forms, and the position of these 
vou. xv.— Part v. No. 6.—December, 1900. Z 
