1886.] MR. F. E. BEDDARD ON A LARGE EARTHWORM. 173 



of the peiiial setae agree with those of A. ungulutus as described by 

 Perrier '. 



Tlie ovaries and oviducts I liave been unable to find. 



There are two pairs of spermatheccB situated in segments 8 and 9 ; 

 each consists of a spherical thin-walled sac communicating with the 

 exterior by a long stout-walled duct which is often curved (see 

 figs. 5 and 6, cp) ; at the upper extremity of the duct, where it unites 

 with the sac, it becomes somewhat bulged out on one side, though 

 there is hardly so marked a diverticulum as is figured by Perrier 

 in A. ungulatus '. 



A very characteristic and remarkable series of structures now 

 remain to be described, before concluding the account of the genera- 

 tive system. In describing the external characters attention was drawn 

 to the modification of the lowermost pair of setas in segment 8; the 

 ordinary setae, at any rate on one side of the body, have disappeared 

 and are replaced by a conspicuous orifice through which protrude 

 one or more stout long setoe, which appear on a naked-eye inspection 

 to be very similar to those which project through the male genera- 

 tive pores. Fig. 5 of PI. XIX. represents tlie internal structures 

 which correspond to these peculiarly modified setse; the latter are con- 

 tained in a thin-walled transparent sac (s) precisely as are the genital 

 setse ; on either side of this sac is a long somewhat sausage-shaped 

 glandular body {g-b), which communicates by a slender duct with 

 the orifice through which the setse project on to the exterior. The 

 presence of these glands renders the whole structure more similar 

 still to the male generative pore, except that the "prostates" are 

 paired. These structures were only present on the left side of the 

 body in one specimen ; in two others they were only developed on 

 the right side; in two other specimens both immature, without a 

 clitellum, these structures were entirely absent on both sides of 

 the body ; in a sixth specimen, which was also immature, with the 

 clitellum undeveloped and with very minute spermathecse, the bundle 

 of modified setse was plainly visible on both sides of the body, but 

 without its accessory glands ; in the seventh specimen, also imma- 

 ture, there was no vestige whatever of these structures ; wherever 

 they were absent the setse of the segment were perfectly normal. 



I am not aware that any structures of this kind have been 

 described in any other species of the genus or in any other Earth- 

 worm ; they appear to be novel to the group. At the same time 

 Perrier figures some structures in A. ungulatus^ which may be 

 identical, though the figure and his description are insufficient; the 

 description (p. 92) is as follows: — " Les poches copulatrices sont 

 situees aux anneaux huit et dix. Chacune est nmnie d'un lobe 

 posterieur, assez petit, et n'a pas d'autre appendice. Dans le neuvieme 

 anneau se voient plusieurs sacs glandulaires, separes par une glande 

 multifide." The details are evidently different from the structures 

 that I have just described, but the similarity of position, between 

 the two spermathecse, renders it possible that they are identical ; in 



1 Loc. cit. p. 90, pi. ii. figs. 21, 22. 



^ Loc. cit pi. ii. fig. 20. ^ Loo. elf. pi. ii. fig. 18, .v. 



