118 PROFESSOR OWEN ON THE ANATOMY OF 
lines : this claw is curved, with its inner border and convex extremity trenchant ; it can 
be bent so as to touch the carpal pad, but cannot be extended so as to bring the end of 
the claw in the line of the digit ; its movements are limited to one plane. The ungual 
phalanx of the third digit, ‘ medius,’ is free: the length of the claw following the 
convex curvature is four inches, its greatest basal breadth nine lines ; its basal circum- 
ference two inches six lines ; its under surface is flat, bounded by trenchant borders ; its 
extremity is more pointed than that of the index ; it can be bent so as to touch the carpal 
pad, but forms an obtuse angle downwards with the digit in a state of greatest extension. 
The second and ungual phalanges of the fourth digit are free; they are compressed, 
and terminate in a straight compressed pointed claw six lines in length; the joint of 
the last phalanx permits a slight extent of flexion and extension, and of free movement 
from side to side. A semilunar notch on the outside of the base of the liberated por- 
tion of the fourth digit indicates the extremity of the abortive fifth digit. This, with 
the metacarpal and proximal phalanx of the fourth, supports a convex callous pad at the 
outer and under part of the sole, two inches by one inch and a half in extent. The corium 
of this digital pad is developed into a number of large, low, obtuse papilla, perforated 
or notched at the apex and covered by larger and more complex papille of the thickened 
cuticle. The carpal pad is smaller, of a kidney shape, one inch five lines in breadth. 
The part of the foot which receives the superincumbent weight in ordinary progression 
is the digital pad, and the outer side of the free portion of the compressed fourth digit. 
The length of the sole of the hind-foot is five inches. ‘The naked part begins about 
one inch in advance of the prominence of the calcaneum: the breadth of the sole at the 
base of the digits is two inches six lines, and there is a callosity at the inner margin one 
and a half inch behind the innermost digit, upon which the supplementary tarsal ossicle 
rests. Each of the five digits has the second as well as the ungual phalanges liberated, 
and each supports a curved, obtuse claw, about one inch in length, but somewhat longer 
on the middle digit and shorter on the outer digit. The three middle digits project 
the furthest, and their claws terminate at the same line: the outermost or fifth digit 
ends a little short of these, and the innermost digit is still shorter. 
The integument of the Great Anteater has something of a pachydermal character ; 
and although there is not any extensively diffused panniculus carnosus, there are some 
well-developed dermal muscles having attachments to parts of the endo-skeleton. 
The corium between the rami of the lower jaw is one line in thickness, and increases 
as the skin approaches the sternum to a thickness of three lines, which is its average 
thickness over the back and sides of the trunk; it becomes rather thinner where it 
covers the abdomen, and upon the limbs. 
The skin is connected to the subjacent muscles, by a thin layer of tough elastic 
cellular substance, along the under part of the neck; but, near the sternum, and over 
the fore part of the sternum, the lobules of an immense salivary gland, resembling 
fat-lobules at first sight, were found interposed. Pressure upon this glandular mass 
(Pl. XXXVII. fig. 1, a, a')—the size, shape, and disposition of which will afterward be 
