092 MKSSRS, B. C. A. WINDLE AXD F. G. PARSONS ON [DeC. 19, 



Family OiiycTEROPODiDTE, 



(G-alton, VIII.) 

 (Humphry, IX.) 

 (Cuvier &' Laurillard, XVII.) 



Ectoqluteiis {Gluteus maximus) and fJaudo-femoralis {Agitator 

 crmdce). — In the Bradypodidce the ectogluteus rises by fascia from 

 the sacral and caudal spines ; according to some authors from the 

 crest of the ilium also. As its origin is fascial, there is clearly con- 

 siderable scope for diversity of opinion and description on this 

 point. 



The insertion is into the shaft of the femur from just below the 

 great trochanter to the middle or rather lower. No separate 

 caudo-femoralis has been described in these animals, except in 

 Cuvier & Laurillard's specimen (6), in which it was very small. It 

 seems quite probable that, in the other specimens of which de- 

 scriptions exist, it is fused with the ectogluteus. This hypothesis 

 is supported to a certain extent by the fact that in our specimen 

 of Cholcepus (8) the muscle had a double insertion, the anterior 

 fibres passing to just below the great trochanter, thus obtaining 

 the normal insertion of the ectogluteus in mammals, whilst the 

 posterior fibres reached the middle and lower thirds of the femur. 

 Among the Myrmecophagidce the ectogluteus and caudo-femoralis 

 can usually be separated from one another, the former passing to 

 the upper part of the femur and to the fascia lata, the latter to the 

 lower part of the bone. This wa" certainly the case in Mynneco- 

 phaga (11) and Tamandua (1-4, 16) and, to a less extent, in Cyclo- 

 </mrt« (21), though most of the dissectors of this animal do not 

 seem to have recognized the caudo-femoralis as a separate muscle. 

 In the Dasypodida: and Manidce both the ectogluteus and caudo- 

 femorahs are present, though usually closely united. In C'hlamy- 

 dophonis (28 a) the latter rose separately from the sphteroma end 

 of the lower sphseroma support. In the OryctL'rojyodidm the ecto- 

 gluteus is inserted into just below the middle of the femur, and the 

 caudo-femoi'alis into the lower end (37). 



Tensor fascice femoris and ilio-tihialis {Sartorius). — In the 

 Bradypodido' the ilio-tibialis is a well-mnrked muscle rising from 

 the crest of the ilium and passing to the inner side of the head o£ 

 the tibia. This applies to Brady pus (1. 4, 5, 6) and Cholcepus 

 (8, 9). The muscular fibres rising external to this and in the same 

 plane, instead of going to the fascia lata, accompany the ectogluteus 

 to the outer surface of the shaft of the femur, but we are of 

 opinion that they represent the tensor fascii« femoris. 



In the Myrmecophagida the ilio-tibialis is distinct and has the 

 human attachments in Myrmecopihaga (11) and Cydothurus (17,21), 

 In Tamandua (14) and Cydothurus (20) it rises from the tendon 

 of the psoas magnus or parvus. The tensor fascise femoris is in- 

 separable from the ectogluteus. In the Dasypodida' both muscles 

 are present, but whereas the ilio-tibialis is a delicate muscle in 

 Dasypus (22, 23, 24), it is thick and fleshy in Chlamydophorus 



