1894.] 



NEW GENEEA OF EARTHWORMS, 



389 



makes up the greater part of the calciferous gland of GordiodrUus. 

 Through this tissue pass numeroas small blood-vessels of equal 

 calibre, which radiate out from the top of the gland but run 

 parallel through its substance. A single diyerticulum from the 

 CEsophagus dips down into it, but appears to end csecally and not 

 to be continued on into a few inti'acellular tubes such as I have 

 described in Gordiodrilus. In the xth and xith segments the 

 oesophagus to some extent retains the structure of the calciferous 

 gland ; outside the lining epithelium of the tube is a granular and 

 nucleated mass which is precisely like the glandular mass of the 

 calciferous gland ; outside of this are the muscular layers and the 

 peritoneal covering of the gut ; it seems therefore probable that 

 the granular tissue of the calciferous gland is of h3'podermic origin 

 and is not formed out of the modification of the peritoneum. The 

 intestine begins in the xiith segment. The nephridia commence 

 in segment v. ; they have no muscular end sac. The last heart is 

 in segment xi. 



The reproductive organs are constituted upon the plan of those 

 of Gordiodrilus, but there are differences of detail. The testes and 

 sperm-duct funnels are in x. and xi. The two sperm-ducts run 

 side by side along the ventral body-wall until about the xvith 

 segment ; after this they get to lie in the body-cavity and are to 

 some extent coiled ; they pass back beyond the point where they 

 open on to the exterior, which is no doubt correlated with the 

 protrusible termination of the efferent apparatus. When the 

 worm is dissected the most obvious part — indeed practically the 

 only part to be seen on account of the small size of the worm — of 

 the efferent apparatus is a pair of oval or pear-shaped sacs ; these 



Fig. 3. 



Nannodrilus. 

 Male efferent apparatus. 



a. Spermiducal glands. 



b. Bursa copulatrix. 



c. Penis. 



Proc. Zool. Soc— 1894, No. XXVI. 



d. Orifice of one of the spermi- 

 ducal glands. 

 v.d. Sperm-ducts. 



26 



