218 BR. R. W. SHUFELDT ON [-'^P'"' 1» 



The posterior uasal fossae are elongo-pear-shaped apertures with 

 the bulbous ends directed backwards, while either palatiue foramen 

 is of an oval outline and of no great size. The inferior temporal 

 fossa is ca[)acious, and a firm thin plate of cartilage standing verti- 

 cally in the median plane divides the orbital cavities internally. 

 This is the interorbital septum. 



Of fair extent, the basis cranii is nearly a horizontal surface, 

 showing bnt a very slight general concavity over it. This is at 

 variance with such a form as lyuanu tuberculatu, where the area 

 to which we refer is considerably concaved. In a previous paragraph 

 we have already sufficiently referred to the tympano-eustachiau fossa 

 and the characters of the columella auris. 



Whatever may be the condition of the parietal ossifications in the 

 very young Heluderma, they are in the adult reptile re|)rescnted 

 solely by a solidly ossified and dense plate of bone. This bifurcates 

 behind, and either limb is directed backwards and outwards and 

 slightly downwards to articulate with the squamosal of tlie same side 

 as ■well as with the corresponding |)arotic process. Viewed from 

 above the anterior margin of the parietal plate is represented by a 

 finely serrated transverse line ; the superior surface of the bone is 

 neaily horizontal and usually supports a group of the ossified dermal 

 tubercles, wiiich have fused with it. Near the middle of its ventral 

 surface is seen a small pit, wliich it would seem is situated too far 

 back to represent the vestige of the parietal foramen. It by no means 

 pierces the bone. At some distance within its external free margin, 

 on eitlier side, this bone develops a longitudinal ridge. This is most 

 conspicuous near its middle, and resting here against its outer aspect 

 are the upper ends of the columella and the prootic. 



In old specimens of this Reptile, the frontal hones are indistiuguish- 

 ably fused together, and \i\)0\\ a superior aspect of the skull not a 

 trace of the niediMu suture that originally stood betweeu them can 

 be made out. And even within the cranium it is hardly to be 

 discerned at all. By a straight transverse coronal suture, this frontal 

 bone articulates posteriorly with the parietal ; while we have already 

 mentioned the fact as to how it is ])revented from participating in 

 the formation of tlie orbital ]ieriphery by the meeting of the pre- 

 and j)OstiVontal elements. Posteriorly, these united frontals are 

 almost entirely masked from our view by the layer of fused and 

 ossified dermal tubercles that overlie the entire fore part of the 

 skull. One never meets with skulls of old individuals of H. sus- 

 pectuiit as free from this feature, nor with the naso-frontal and 

 fronto-parietal .sutures anything like as clearly defined as is seen in 

 the skull of //. horridum which is figured for us by INIivart 

 (Encycl. Brit. Dth ed. vol. xx. p. 451, fig. 12 /.). Ventrally, the 

 frontals of the skull of our present subject offer us a peculiar cha- 

 racter. Opposite the orbits each one sends downwards and inwards a 

 broad and curved plate of bone which mesially meets and fuses with 

 a corresponding plate coming from the bone of the other side. This 

 arrangement gives ris^e to a transverse osseous bridge, and the large 

 mesial foramen it assi&ts to form has passing through it certain 



