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glabrous, ae angled, psa ries by a narrow or broad 
callus-ring; pappus in 2-3 serie s, capillary, its bristles all equal or 
nearly so, at least not in two unequal series. Shrubs with pubescent 
or tomentose stems and large rugose obovate leaves, usually scabrous 
above and tomentose beneath. 
Type, Vernonia Letboldiana Sch.-Bip. 
Four species, natives of southern Mexico. Distinguished from 
Vernonia primarily by the structure of the pappus, the genus has 
also avery different habit, due to the huge crowded heads and large 
broadly obovate leaves. The callus-ring surmounting the achenes 
becomes obscure at maturity. 
Leaves thin ; heads peduncled ; scales glabrous or nearly so, acute to obtuse, 
incipal ones 
Leaves green beneath, — strigose. 1. L. Salvinae. 
Leaves cano-tomentose beneath. 2. L. mexicana. 
Leaves thick ; heads nearly ee scales pubescent or pares regularly 
taperi ng to a long point, the principal ones 2 mm. wi 
Scales prominently carinately nerved ; achenes tipped wih a narrow 
callus rin 3. L. Leiboldiana. 
Scales not ey nerved ; achenes usually a a broad 
callus 4. L. serrata, 
1. Leiboldia Salvinae (Hemsley) nom. nov. 
Vernonia Salvinae Hemsley, Biol. Cent. Am. Bot. 2: 73. 1881. 
Herbaceous or shrubby; ieee ta branches li Seales -tomen- 
tose, becoming glabrate with a cauline leaves membranous, 
Sede cine to narrowly obovate, 15-25 cm 4-7 cm. 
wide, short-acuminate, narrowed below into short ‘petioles, rather 
remotely serrate with low callous teeth, pinnately veined, puberu- 
lent above, especially along the midvein, strigose-hirsute ‘beneath ; 
heads 1-5, on peduncles 3-7 cm. long: involucre broadly campanu- 
late, 15 mm. high; scales thin, ‘purple, somewhat spreading at the 
tip, ovate to oblong, rounded to subacute at the apex, or the inner 
most acuminate, puberulent and glandular, especially near the tips; 
achenes sharply 4-5- ona with a prominent callus-ring; pappus 
tawny, about 7 m 
Type locality: Guatemala: 
Distribution: Chiapas, Mexico, Melson 3747, to Guatemala 
and Costa Rica, Wercklé rr60¢4. 
. Salvinae is a little-known species poorly represented in 
herbaria. Hemsley made no mention of the structure of the pappus, 
although he described the tetragonal achenes. Its supposed rela- 
tionship to Vernonta Alamani DC. is merely one of habit, the two 
differing completely in the structure of pappus and achenes. 
