( 334) 
from near the bottom of the cup give rise to the gynoecium. 
They grow in toward the center until they meet and then be- 
come prolonged upward to form the style and stigma. From 
one of these outgrowths a small elevation may be early dis- 
tinguished, which marks the rudiment of the ovule. This 
elevation bends down toward the base of the ovary and soon 
fills the cavity of the ovary before there is any differentiation 
of integuments. By the time the cavity of the ovary is filled 
the young ovule is a downward projecting lobe and about 
the time that the integuments are first distinguishable it 
begins the turning which finally brings it to the anatropous 
position. The turning takes place toward the side on which 
the ovule had its origin, the apex of the nucellus being 
turned underneath and finally brought up toward the pla- 
centa, so that its final position is anatropous, with the micro- 
pyle directed toward the style. The single ovule entirely 
fills the ovarian cavity, but it is at all points separate from 
the wall except at the placenta. 
The development of AZicrampel:s presents some very inter- 
esting differences from the foregoing types. The pistillate 
flowers are borne singly and each normal ovary contains 
four seeds. In the mature fruit the micropyles of the ovules 
are directed toward the base of the ovary, the opposite, in 
this respect, of Szcyos. The earlier stages in the develop- 
ment of the ovary resemble those of Szcyos, in the manner of 
the formation of the cup and the development of the two 
apparent rings inside the cup. The upper ring grows upward 
forming style and stigma, as in S7cyos. The lower ring be- 
comes modified in a very different way. The ovary here is 
evidently composed of two carpels and the inflexed borders 
of these carpels as usual become the placentae. What 
appears to be a ring is not actually such but opposite eleva- 
tions which mark the edges of the carpels and which are more 
or less convergent horizontally. These elevations appoach 
from opposite sides and flatten against each other at the center, 
so that two lobes are formed each provided with two wings. 
On the lower portion of each wing an ovule arises in the 
