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usual manner. The apex of the nucellus is at first directed 
downward but gradually turns toward the axis of the ovary. 
The turning continues so that the micropyle is at one time 
directed toward the axis, later upward, and finally downward 
again by continuous turning in the same direction. As in 
Szcyos the cavity of the ovary is very early filled by the grow- 
ing tissue and the ovules, but here also the stylar canal is 
maintained. While the turning of the ovule is taking place 
a torsion of the funicle is also going on so that the raphe in- 
stead of lying on that side of the ovule which is toward the 
median line of the carpel, is on the side toward the line of 
union of the carpels. The placentation, which is at first ap- 
parently basal and later central, is in reality parietal and es- 
sentially similar to that of the other types already considered. 
Cyclanthereae 
Near the growing point of the shoot, in Cyclanthera, there 
arise lateral protuberances (leaf-rudiments), and in the axils 
of these outgrowths other lobes appear which are the prim- 
ordia of the inflorescences. ‘The division of the axillary lobe 
into lesser lobes is soon apparent. One of these becomes a 
pistillate, the others all staminate flowers. The difference 
between the two can early be recognized. Both at first are 
flattened across the apex, but soon the staminate flowers are 
marked by a convexity within the circle of corolla lobes, while 
the pistillate flowers become more concave. In the pistillate 
flower this concavity deepens and the placenta appears as a 
broad ridge inside running parallel with the axis of the 
flower. This placental ridge soon fills the cavity and bears 
from ten to fifteen ovules in two lateral rows. The stylar 
canal, easily discernible in the earlier stages, is practically 
obliterated at the time of the fertilization. At this time the 
ovules stand almost perpendicular to the axis of the ovary 
with the micropyle directed outward near the line of union 
of the carpel-borders. 
