(341) 
more than its grosser morphological features. It consists 
(jig. 27) of an egg-cell, two synergids and two polar nuclei, 
at a time when the pollen-tube can be detected in the tissue 
just outside the micropyle. The synergids are long, narrow 
and vacuolated in their lower portions. The egg-cell is rela- 
tively large, but consists of only a thin protoplasmic sac with 
the nucleus at the lower end. The polar nuclei are largest 
and surrounded by a dense mass of protoplasm. No antip- 
odals could be found. 
Several stages in the development of the embryos were 
observed. The earliest walls found are transverse, but these 
are soon succeeded by longitudinal divisions (figs. 28-32), 
first in the distal cell of the series, and later in the short or 
rudimentary suspensor. The cells of the suspensor are never 
many, and the few that are formed are soon disorganized 
(figs. 30, 31). The first cleavage of the proembryos are 
quite regular (fg. 37), but the subsequent divisions are in no 
definite order, and a globular mass of cells is soon formed 
(fig. 32) in which no differentiation is apparent except that 
the epidermis is clearly defined and remains so. A beak-like 
prolongation in the proximal portion indicates the slight sus- 
pensor, which is not always connected with the wall of the 
embryo-sac cavity ; the embryos usually appearing to be free 
in the micropylar end of the cavity. ‘The spherical form of 
the embryos is maintained until they reach a size visible 
through an ordinary hand-lens. An elongation of the mass 
then begins and the cotyledons soon appear as two lobes at 
the distal extremity (7g. 33). In the next later stage ob- 
served the cotyledons had assumed the proportions indicated 
by figure 3g. Still no differentiation of periblem and ple- 
rome could be detected, but the initial stages in the formation 
of the root-cap were visible. 
The endosperm begins to form early after fertilization. 
The complete fusion of the polar nuclei was not observed, 
but if it occurs it must be at about the time of fertilization. 
By the time the first cleavage of the embryo has been com- 
pleted, the endosperm has formed a thin sac which rapidly 
