44 MEMOIRS OF THE NEW YORK BOTANICAL GARDEN 
cylinder has grown larger and shows a concave depression on its 
upper surface. The bundles at this stage begin to separate from 
each other and, contrary to the conditions found in Sequoia, diverge 
widely into the very much flattened peltate portion of the cone scale. 
So quickly does the expansion of the narrow stalk into the flattened 
terminal shield-portion take place, that it 15 almost impossible to 
secure a section showing the transition from the one to the other. 
The small bundles arrange themselves around the margins of the 
disk and become completely surrounded with transfusion tissue, а 
feature of marked contrast to the scale bundles in Sequoia and at the 
same time one which indicates a strong affinity with the Araucarineae. 
In Sequoia, as has been pointed out by the junior author,*” there are in 
the upper portion of the bundles of the cone scales lateral wings of 
transfusion tissue connecting the bundles with one another. In fact in 
the cone scales of the Sequoiineae and Cupressineae there is present 
the same type of transfusion tissue which is shown in the leaf-bundle 
of Cunninghamia sinensis, as figured by De Bary.*! In this type of 
transfusion tissue the tracheidal cells originate from the flanks of 
the bundles and do not encircle the whole strand as in the vegetative 
leaves of the Abietineae or envelop the phloem part of the strand 
as is often the case in the bundles of the cone scales of living repre- 
sentatives of the Araucarineae. It has not been possible to trace 
any of the strands of the peltate portion of the cone scales under 
discussion into cicatrices of ovules, so that the attachment of the 
ovules in this case was probably on the stalk of the peltate scale near 
the axis of the cone. This view of the matter is strengthened by 
the discovery of one immature and carbonized cone composed of 
peltate scales, in which there are the remains of four ovules on the 
peduncular portion of each scale. Unfortunately the specimen is 
completely carbonized and cannot be sectioned to discover its pos- 
sible affinities with the isolated mature scales described above. Figure 
3, Pl. 25, shows a transverse section, X 100, of one of the numerous 
small bundles found about the margin of the disk in the mature 
scales under discussion. It may be clearly made out that there is a 
radial disposition of the transfusion tissue around the bundle, which 
appears as a cluster of smaller cells in the center of the larger trans- 
fusion elements. 
“Mem. Boston Soc. Nat. Hist. 5: 449. pl. 70. f. 2 
= a Anatomy of the Vegetative Organs of d Phanerogams and Ferns, 
fg. 183, p 
