MUSEUM OF COMPAKATIVE ZOOLOGY. 357 



evidence of this is found in the arrangement of pigment granules, fat 

 drops, or (in the younger larval stages) remnants of the yolk granules, 

 which in the quiet cell lie grouped uniformly around the nucleus or 

 are quite irregularly distributed. " Wenn aber der Kern in die erste 

 Theilungsphase tritt, hahen sie sich zu zwei Gruppen geordnet.'''' * These 

 groups in oval cells usually correspond with the poles of the longer axis, 

 the plane of division with the shorter axis ; in some rare cases, however, 

 this relation is reversed. When the division-axis of the cell chances to 

 be very oblique to the plane of the object stage, the grouping of these 

 " polar granules " is seen less in prohle, and then exhibits a distinct 

 radial arrangement (loc. cit., Taf. XVI. Fig. 6 a). 



Phase 2. The loose glomerate or basket form t of the mother nu- 

 cleus. This more open-meshed condition implies a diminution in the 

 number of the filaments, or, more probably, a diminution in the total 

 length of the filament, and is accompanied by a corresponding increase 

 in thickness, so that the volume of stainable substance remains the 

 same as in the first phase. This thickening, it is believed, is not brought 

 about by a direct fusion of neighboring filaments, — since no filaments 

 are found which are in part of their course as slender as in the first 

 phase, and in the adjoining part twice as thick, — but is due to a slow 

 process, shortening the filaments, and at the same time making them 

 correspondingly thicker, as in the contraction of a muscle fibre. This 



of overlooking early changes in the protoplasm is greatly increased, and, what is of 

 more importance, that these tissue-cells can hardly be claimed to be favorable objects 

 for determining when the earliest appearance of a dicentric arrangement in the proto- 

 plasm takes place. I will not, however, insist here on the legitimacy of what I have 

 only ventured to call attention to in another connection as seeming worthy of more 

 careful attention before the initiative and controlling influence in cell division is 

 definitely — not to say exclusively — assigned to the nucleus (or nuclear substance)- 



* The words here italicized are not so in the original. 



t The basket form here spoken of is not identical with a condition of the nucleus 

 thus named by Mayzel and Eberth. The structure intended by them is called by 

 Flemming cask- or half-cask-shaped. It is only the staves of the cask and the ribs of 

 the basket which are represented in these figures. Flemming recognizes the inapt- 

 ness of the expression *' basket-form " (previously employed by him) to represent this 

 glomerate condition for the following reason : the glomerule of tortuous filaments, 

 although it is more compact at the periphery than in the centre, is not simply limited 

 to bounding a cavity. On this account he prefers the name glomerule (Knauel), even 

 though this is objectionable as implying that the thread is wound about a definite 

 centre, which is really not the case. 



If ravelled 5'^arn be simply balled together without winding, the condition of the 

 I tortuous filaments will be very well reproduced ; it would only remain to make the 

 mass somewhat more compact at the periphery than in the centre. 



