DECEMBER 22, 1899. ] 
T. proriger, making 50. Thirty-eight (38) 
elements are preserved in the left fore paddle 
or manus and 44 are inserted in our res- 
toration, or 5 metacarpals and 39 phalanges. 
In the hind paddle, or pes, 33 metatarsals 
and phalanges are preserved on the left 
side (including an isolated phalanx which 
lies above the 50th caudal). 
The phalangeal formula is estimated as 
follows : 
MANUS. PEs. 
Digit I. 5 5 
THE 8 8 
III. 8 8 
IV. 9 & 
Vv. 9 6 
T. dyspelor. T. dyspelor. 
It is apparent, so far as we can judge 
from this specimen, thatin TY. dyspelor the 
phalanges are less numerous than in 7. 
proriger. ‘ 
2. A second characteristic is the marked 
broadening and shortening of the 5th metapodial 
in both manus and pes, but especially in 
the pes. The carpus and still more the 
tarsus, on the postaxial (ulnar and _ fibular) 
sides are abbreviated. The result is that 
the 5th digit is drawn towards the body ; 
its elements and joints alternate with those 
in the I.-IV. digits; as a whole it is set wide 
apart. Williston has recently shown that 
the epidermal fin web conforms in its con- 
tours to this peculiarity. 
3. A third characteristic is the alternation 
of the joints in the Ist and Sth digits with those 
in digits2, 3,4. The pes further agrees with 
the manus in the expansion of the proximal 
part of metapodial I., and the shortening or 
drawing up of the first finger, whereby the 
middle points of the phalanges of Digit I. 
come opposite the joints of the phalanges in 
Digits IJ., I1I., 1V., thus greatly strength- 
ening the paddle asa whole. A _ similar 
adaptation by alternation of the phalangeal 
joints is observed in some of the Plesiosaurs, 
in which it is carried to an extreme, for 
the phalanges of all the digits alternate. 
- SCIENCE, 
923. 
This specimen affords an exceptionally 
favorable opportunity for a restoration of 
the skeleton. This interesting work has 
been accomplished by cooperation. Dr. 
W. D. Matthew kindly undertook a nat- 
ural-size drawing of the entire animal, suc- 
ceeding especially in rearranging the ver- 
tebral column and skull. Mr. Horsfall 
completed the details of the skull by care- 
ful measurement and comparison with the 
drawings of Merriam and Williston. Dr. 
McGregor and the writer restored the pad- 
dles and the sternum. 
The drawing is upon a one-eighth scale. 
There was probably a small rib upon the 
third and fourth cervicals which has not been 
indicated. The cervical intercentra are re- 
stored from a fine specimen of Platecarpus. 
One of the most important features of 
the restoration sprang from the discovery 
that the cartilaginous ribs of the left side 
are practically in their normal relations. 
This fact enabled us to locate definitely the 
lower end of the ten true ribs, the sternum, 
the epicoracoids, and at the same time fix 
the position of the fore paddle with refer- 
ence to the skull. 
As above noted, the ribs were found to 
resemble those of Sphenodon much more 
closely than those of Varanus. They are 
thus given in the restoration the angle, 
position, and foreshortening characteristic 
of Sphenodon, as the narrow anterior part of 
the chest expands into the broader walls of 
the abdomen. The ribs in the plate are 
perhaps a shade too heavy. 
The upward curvature of the tail is de- 
signed exactly as the vertebree lie in the 
specimen, for the reasons already discussed. 
RESTORATION OF THE ANIMAL. 
In the restoration of the animal, Mr. 
Charles Knight has taken advantage of all 
the information afforded by Professor Will- 
iston’s collections and descriptions, and of 
our detailed study of this fine specimen. 
