ANATOJiy OF TUB GOUT-FLY OF BARLEY. 807 



;mil back ou each side to the posterior region of the luetathoracic 

 soginent. The peduncles of the wing discs reach the hypoderni 

 at a point immediately opposite this lissure. The vertical splits 

 Avluch pass up and down the lateral regions of the pnparium 

 from the posterior ends of the lateral splits do not appear to be 

 similarly marked out in the third instar larva. 



Gonadial Rudiments. 



The rudiments of the ovaries or testes lie one on either side of 

 the fifth abdominal segment in its posterior region. They are 

 solid oval masses of cells with the long axis parallel to the long 

 axis of the body, and are attached to the neighbouring fat body. 

 As I have not so far been able to distinguish a larval ovary from 

 ix. larval testis, it would be quite useless to give any account of 

 the structure of the gonadial rudiments. 



Muscular System. 



The details of the muscular system have not been worked out. 

 The arrangement of the muscles probably approximates very 

 closely to their arrangement in Musca domestica which is de- 

 scribed by Hewitt (4). 



0. Other Immature Stages of Ciilorops, 

 21ie Puparium (text-fig. 12). 



Length 5-4 mm,; maximum breadth l-y mm. Colour ex- 

 tremely variable; all shades between pale yellow to very dark 

 brown. Thoracic segments slightly dorso-ventrally compressed, 

 much wrinkled transversely and obliquely, always darker coloured 

 than the abdominal segments; apices of wrinkles usually appear- 

 ing dark brown or black, the spaces in between beino- lighter. 

 The last two abdominal .segments are also considerably wrfnkled 

 and darker coloured than the others. The other abdominal 

 segments are also transversely wrinkled, but the wrinkles are not 

 pronounced or close together and are not darkened. The larval 

 cephalo-pharyngeal skeleton lies pressed against the ventral wall 

 in the anterior end of the puparium. The larval head is com- 

 pletely withdrawn within the prothorax and is quite invisible. 

 The larval stigmata stand out clearly. There are no pupal dorsal 

 respiratory trumpets projecting through the dorsal wall of the 

 pupaiium {cf. Keilin, 1917). 



For the exit of the imagines the anterior end of the puparium 

 splits into dorsal and ventral halves by means of a horizontal 

 split running round its anterior end and backwards alou'' each 

 side as far as tlio anterior region of the first abdominal se"ment. 

 From each posterior end of this split a dorsal and a venti-al split 

 extends round the anterior end of the first abdominal se^^ment. 

 The extent of these transverse splits is very variable ; sometimes 

 they ai-e very short, sometimes the dorsal ones meet in the mid- 

 53* 



