888 
Hemitelia by Bonnier (J. ¢.), and contain 
relatively much more protoplasm than the 
contiguous cells, large nuclei and chloro- 
plasts. The cytoplasm is vacuolated and 
the cell walls are thin. The cells stand in 
contrast with the ground parenchyma on 
account of their smaller size, their diame- 
ters being between the ratio of 1 to 3 and 1 
to 4. The cells near the epidermis are 
larger than those lying deeper in the gland. 
In making the foregoing statement, we dif- 
fer from Figdor, who says ‘‘ The single ele- 
ments of the gland are about the size of 
the cells of the fundamental parenchyma.” 
Frequent small intercellular spaces occur. 
These connect with each other and finally 
with the substomatic spaces which are of 
considerable size (Fig. 2). Through these, 
therefore, the secretion may findits way to 
the surface of the gland. ‘Bonnier, in the 
paper above cited, makes no mention of the 
substomatic spaces in the special treatment 
of the ferns. Further on, however (p. 
151), he makes a general statement to the 
effect that he had established the fact that, 
in the cases of nectaries with stomata, these 
are either without a substomatic space or 
have only very small ones. His figure 
representing the nectary of Hemitelia obtusa 
(on his Pl. 1, Fig. 9) certainly bears out 
his general statement. We cannot, how- 
ever, regard this figure as satisfactory. No 
attempt was made to delineate the stomata, 
except in very schematic fashion, on ac- 
count of the small size of the drawing, for 
whichreason also the possibility of represent- 
ing substomatic and intercellular spaces was 
very much lessened. The presence of a 
large substomatic space, however, may not 
be regarded as of any importance in the 
economy of such an organ as a nectary, in 
which the movement of the fluid is the 
important and characteristic feature. 
Passing on to consider the relation of the 
nectary to the vascular tissue, we notice in 
tie first place the distribution of bundles in 
SCIENCE. ; 
[N. S. Von. XIII. No. 336. 
the immediate vicinity of the former. Fig. 
4 shows, in diagrammatic form, this re- 
lation, from which there appears to be little 
variation to be noted beyond. A broad 
Fic. 4. Diagram showing ‘the relations of the 
vascular tissue and gland. The dotted outline shows 
the extent of the latter. c, limit of the cribral part of 
the bundle ; e, endodermis. The pericycle is included 
between these. 
stele* (st) divides at the lower angle of the 
nectary, one branch (p’) running along one 
side of the gland and thence into the pinna, 
the other and larger passing along the other 
border of the gland, broadening as it goes 
into a vascular plate which consists of a 
dense complex of short, irregular wood ele- 
ments, well-developed cribral cells and a 
several-layered pericycle, to be described 
below. 
From this plate of vascular tissue, which 
lies beneath and somewhat obliquely across 
the nectary, pass forward two branches, p 
and 7, which go into the pinna and rachis, 
respectively. Sometimes four branches 
arise by the splitting of these two, but the 
general character of the arrangement is 
quite constant. Sometimes a small stele 
runs into the complex, as shown ata, Fig. 4. 
* The term stele is used here, in the ferns, without 
reference to the question of morphological pro- 
priety. See Jeffrey, E. C., ‘The morphology of the 
central cylinder in the angiosperms,’ Zrans. Can. 
Inst., Vol. 6. 
