16 MR. F. E. BEDDARD ON THE [May 3, 



The azygos veins are to be found on both sides of the body ; 

 but that of the right side is much the larger, and extends back, 

 for the space of about eight ribs, some Httle way beyond the 

 anterior end of the lungs. On the left side there is a very 

 slender azygos, which might readily be missed on account of its 

 delicacy. This latter receives, before entering the left jugular, a 

 branch from the esophagus. In Chamceleon, as I have already 

 pointed out, the oesophageal vein enters the right side of the 

 heart via the azygos vein. 



Lateral Abdominal Veins. — These veins arise at the junction 

 of each root of the anterior abdominal with the vena renalis 

 advehens. They do not arise fi'om the anterior abdominals 

 themselves as is the case with most Lizards. Their course along 

 the body- wall is, however, like that of many other Lizards. On 

 the right side the vein is connected supei'ficially by slender 

 branches with the dorsal parieto-hepatic veins of that side. 

 Anteriorly the veins die away. 



Hepatic Portal System. — As in most, but not all. Lizards, 

 nearly the full complement of hepatic portal vessels is present 

 in Pygopus. It receives blood, that is to say, from the anterior 

 abdominal, from the epigastric, the stomach, and the dorsal parietes. 

 There are only wanting independent vessels from the ventral 

 parietes present in many Lizards and so marked a feature of 

 Chamceleon. 



The epigastric vein appears to be single ; if lateral epigastrics 

 are present they are small — unless, indeed, the vessels which I 

 homolooise with the lateral abdominals of other Lacertilia are 

 really to be looked upon as lateral epigastrics. The single epigas- 

 tric lies to the left of the middle line. It is connected with the 

 anterior abdominal by two branches just behind the liver. It 

 oives off a large number of branches to the liver. I counted 

 eight of these altogether : they extend along the whole of the 

 liver, but do not arise at regular intervals, and are of unequal 

 size. ' Anterior to the liver is a tract of the epigastric which 

 oives off, at any rate, one branch to the vena cava before passing 

 headwards and ending in a way that I have not been able to 

 ascertain. It is noteworthy that here, as in Phelsiima, in both 

 of which there are no marked lateral epigastrics, the median 

 epigastric takes up a portion of the duties of the lateral epigas- 

 trics, and conveys some of the blood from the posterior abdominal 

 region straight to the heart. 



The dorsal parieto-hepatic vessels in this lizard are numerous 

 and large. Altogether three of them enter the liver, and at 

 least two of them are connected, as already mentioned, with the 

 lateral abdominal veins by fine branches running superficially over 

 the parietes. There are also three on the left side, more anteriorly, 

 which communicate with the liver via the stomach. 



There are four gastrohepatic vessels, which arise separately from 

 a continuous longitudinal trunk running along the stomach, which 

 is fed not only from the ve.ssels of the stomach itself, but also by 



