168 ON THE ANATOMY OF LIMICOLINE BIRDS. [May 16, 
side for digits 2 and 4 respectively, except in Glareola and 
Thinocorus, where it is a fan-shaped slip of fasciee common to 
the three digits, with the slightest trace of specialisation into 
tendons on the edges of the fan. 
Peroneus superficialis (with slip to perforated tendon of digit 3), 
Peroneus profundus.—These muscles are present, with one 
exception, in the normal or archecentric condition in all the birds 
on the list. The exception is the peroneus profundus in 
(dicnemus, in which bird it is practically absent, the absence 
being another point in which W@dicnemus differs from the 
Limicole and recalls many of the Gruiformes, such as Otis. 
Flexores perforantes et perforati.—These muscles and tendons, 
including the slip connecting the tendon of digit 3 with the 
corresponding tendon of the perforated flexor, all present a 
practically identical condition, which does not differ in any 
important respect from the condition in the majority of the 
Gruiformes. 
Flexores perforati.cThese muscles in all the birds on my list 
have the usual inter-relations and divide mto tendons for the 
three digits in customary fashion. The muscular mass has three 
heads: of these I have already described the ambiens head, which 
is similar throughout, except that there is no accessory ligament 
from the head of the fibula in Chionis. The external head is 
fleshy in Himantopus; it 1s small and tendinous in Chionis, 
Glareola, Thinocorus, Hydrophasianus, Charadrius, Rhynchea, 
and Gallinago. Itis absent in @dienemus and Scolopax. 
Pleaor profundus and Flexor longus hallucis.—In my communi- 
cation on the Gruiform birds I described various ways in which 
the tendons of these two muscles (which are similar in their 
origin in all the birds on my list) are united with one another 
and distributed to the toes. I suggested that probably the 
most primitive condition was such as is to be found in Hurypyga 
(7, text-fig. 85, VII), where the longus hallucis sends a slip to the 
hallux, and distad of this blends so completely with the profundus 
tendon that each tendon supplies each of the three digits. The 
condition in Chionis resembles this closely, except that, as in 
Rhinochetus, the hallucis tendon, after giving off its slip to the 
toe, is not so markedly spread out for the other toes. Mdicnemus 
shows a state practically identical with that of Otis; there is no 
great toe, and therefore no slip to it; the spreading out of the 
junction of the hallucis tendon with the profundus tendon has 
become obliterated. 
In Hydrophasianus the condition is exactly as in Hurypyga, 
except that, although there is a long great toe, there is no slip to 
it. In Rhynchea the condition is also the primitive one, except 
that the slip to the great toe comes off a considerable distance 
above the branching of the conjoined main tendons for the three 
other digits. In Scolopax the condition is similar to that in 
Rhynchea, but although there is a small great toe there is no 
slip to it, and the long junction of the two tendons is ossified. 
