464 MR. F. E. BEDDARD ON THE [ Noy. 28, 
as is also often the case with the Lacertilia, are separated by the 
dorsal mesentery. 
The afferent renals are, as is the case with Lizards, derived 
from two sources: the caudal vein divides into the two veins of 
Jacobson and there is also a system of vessels derived from the 
hind limbs and from the parieties in that neighbourhood. I 
traced the veins of Jacobson for some way into the substance of 
the kidney. It appeared to me that they did not directly join 
the anterior abdominal vein ; and in any case it seems clear that 
instead of there being a branch superficial to the kidney which 
joins the ischiadic afferent renal system as in other Lizards (for 
instance, in Lacerta, as figured by Hochstetter *), there is at most 
a branch which effects such a union running within the substance 
of the kidney. Jam inclined even to think that the union is 
indirect. But in either case there is obviously an approach to 
the condition observable in non-Boine Snakes, where the anterior 
abdominal vein is independent of the caudal vein. It will be 
noted, moreover, that the condition observable in the kidney- 
region of Hatteria is quite remote from that to be noted in the 
Varanide and in the Crocodilia, where the ischiadic, or both 
the ischiadic and caudal, veins are directly continuous with the 
anterior abdominal vein or veins, and merely send branches to 
the kidney. The afferent renal system of Hatteria is, as it were, 
an exaggeration of the typical Lacertilian type. 
It is more particularly the anterior abdominal vein which 
appears to me to show these Ophidian characters, partly matched, 
however, as I shall indicate later, in a legless Lizard, Pygopus 
lepidopus. In Lacertilia, at least as a rule, the conjoined anterior 
abdominal and portal veins enter the left lobe of the liver at or 
quite close to its posterior border. 
In Snakes, on the other hand, there is, at least in some cases, 
a different arrangement. In Hry«, for instance, the portal 
runs along the side of the liver to its anterior end, giving off 
branches at intervals to the liver-substance. In Hatéeria also (see 
text-fig. 60) this is precisely what happens. The anterior abdo- 
minal vein, reinforced by the portal, runs in the membrane which 
connects the stomach with the left lobe of the liver, giving off 
branches at intervals. to the liver-substance and receiving at 
intervals branches from the stomach. ‘Towards the anterior end 
of the liver the conjoined porto-abdominal trunk finally disappears 
in the liver. 
The details of the branching described here in general terms 
can be understood by a reference to the figure (text-fig. 60). 
Pygopus ~ shows an intermediate state of affairs. The main 
branch of the conjoined portal and anterior abdominal veins 
enters the liver near to its posterior extremity, as in Lacertilia 
* Morph. Jahrb. vol. xix. pl. xvi. fig. 12. 
+ “Notes upon the Anatomy of certain Snakes of the Family Boide,” P. Z.S. 
1904, vol. 11. p. 113. 
{ P. Z. S. 1904, vol. ii. p. 17. 
