1905. ] VASCULAR SYSTEM OF CROCODILUS. 467 
sponding vessel in 7%liqua*, which arises on the left side of the 
vertebral column and joins the right parieto-hepatic. In the 
Lacertilia generally the dorsal parieto-hepatic veins are on 
the right side only ; and if there are such vessels on the left they 
either join the right-hand vein before entering the liver, as in 
Tiliqua, or, as in Iguana ty and Anguis f, supply the castric and 
cesophageal networks and thus reach the liver indir ectly. It may 
be said, therefore, so far as present knowledge allows the state- 
ment, that the Crocodilia differ from the Lacertilia in having 
a left as well asa right dorsal parieto-hepatic, entering the liver 
independently into both right and left lobes. 
On the right-hand side of the body the single dorsal hepato- 
parietal trunk is composed of three stout affluents, of which the 
two posterior are joined by a cross anastomosis. The anterior 
vessel is continued forwards superficially and joims, or nearly 
joins, the right azygos. I may remark that on the left side the 
gaps between the several superficially running sections of the 
azygos are more pronounced. 
Finally, the venous system of the liver in Crocodilus acutus 
receives another affluent, which is not, as I believe, represented in 
the Lacertilia. This isa vein which arises from the parietes on 
the right side, in a position intermediate between the dorsal and 
ventral parieto-hepatics. This single vein arises from a longi- 
tudinally running trunk in the parietes which corresponds, as I 
think, to the lateral abdominal vein; it passes straight to the 
liver, which it does not, however, enter independently, but in 
common with the dorsal parieto-hepatic. 
The above-given facts are, as has been already mentioned, not 
in entire accord with previous statements. 
Hochstetter observes § ‘‘ Hine vena hepatica advehens vertebralis 
[=my right dorsal parieto-hepatic] wie bie Lacerta und anderen 
nicht vorkommt.” But his observation refers to Alligator lucius. 
Jacquart ||, who previously studied the venous system of the 
same species (“Caiman & museau de Brochet”), makes no 
particular mention of the vessels which I describe here in 
Crocodilus acutus. He refers to what is possibly the right dorsal 
pavieto-hepatic (figured im fig. 1, pl. ii. 20 d, of his memoir) 
merely as “une petite veine qui contourne le bord tranchant du 
lobe droit du foie”; and the vein in question may be really a 
branch of the ventral parieto-hepatic system. For the possibly 
corresponding vein of the left side (figured by Jacquart in fig. 1, 
pl. iii. 41, of his memoir) no special reference is made in mine 
text. 
Rathke 4, however, under the name of “vena epigastrica 
* Beddard, “Contributions to the Anatomy of the Lacertilia: Pt. I.,” P. Z.S. 
1904, vol. 1. p. 445. 
+ Id. ibid. p. 440. { Morph. JB. vol. xix. p. 473. 
§ Morph. JB. xix. p. 478. || Ann. Sci. Nat. (4) ix. 1858, p. 129 &c. 
@ ‘Untersuchungen tiber die Entwickelung und der K6rperbau der Krokodile’ 
(Braunschweig, 1866), p. 256. The same name is also applied to the abdominal veins ; 
but, I presume, in error for ‘‘ extern.” 
