44 MR. F. E. BEDDARD ON THE ANATOMY [Jail. 15, 



gradations between the extremes to permit of the use in classi- 

 fication of the dimensions of this gland. 



Liver-lobes. — The proportions of the two lobes of the liver* 

 difier markedly in several species of the Ohamseleons reported 

 upon in the present communication. Thus in Gh. immilus the 

 right and left lobes are so nearly equal that only one can be seen 

 with just traces of the other when the visceiu are viewed from the 

 left side. The gall-bladder is partly covered by the extensive left 

 lobe, which, moreover, comes into contact with the stomach. 



The most extremely opposite conditions to these are shown (so 

 far as the material in my hands enables me to say) in Gh. calcarifer. 

 In that Chamseleon the left liver -lobe is very much shorter than 

 the right. When the animal is viewed in the same posture as the 

 last, the left lobe leaves exposed a section of the right lobe as long- 

 as itself, and does not ev^en reach the gall-bladder, which lies on 

 the right lobe not very far from its tip. There is, of course, no 

 contact between the left lobe of the li^•er and the stomach, the 

 ventrally flexed region of which lies considerably behind the end 

 of even the right liver-lobe. 



In Gh. hasiliscits the viscera in question are arranged and have 

 very much the same proportions as in Gh. calcarifer. The same 

 may also be said of the Common Ohamgeleon {Gh. vulgaris) and of 

 Gh. verrucosus t. In Gh. jyarvilohus the two lobes of the liver are 

 approximately equal, and the left lobe completely conceals the 

 gall-bladder when that viscus is viewed from the left side. On 

 the other hand, it, the left lobe, does not come so near to the 

 stomach as in Gh. pumilus. Very much the same description will 

 serve for Gh. dilepis, save for the fact that in this Champeleon the 

 gall-bladder is not so completely hidden by the left lobe as in 

 Gh. jmrvilobus and Gh. jnmnilus. The characters of the liver, 

 therefore, hardly allow of any grouping of the species ; for there 

 are gradations. The alimentary canal does, however, show certain 

 differences which permit of a grouping such as has been already 

 suggested. 



In Gh. calcarifer and the other largei- species the intestine is as 

 well coiled as in other Lacertilia, and when the animal is opened 

 from the side a good deal of the small intestine is seen to lie in a 

 coil with secondary convolutions in front of, i. e. headward of, the 

 pyloric end of the stomach. The stomach, in fact, partly covers 

 a section of the small intestine when the viscera are viewed from 

 the left side. In Gh. pumilus there is only one short bend of 

 duodenum, which lies in front of the stomach. But the opposite 

 extreme to Gh. calcarifer is to be seen in Gh. tceniohronchus. In 

 this small Chanifeleon the end of the stomach is not bent upon 

 itself at all, but is continued back in a sti-aight line to join the 

 intestine, which is but little coiled upon itself. Moreover, the 

 whole length of the intestine lies completely behind the stomach. 



* The liver itself is very compact in these reptiles, unlobulated, and with very 

 fii'm outlines. 



t I could see no gall-bladder in this species. 



